chr3 : 190,325,950 – 190,326,598
648 bp 76 TFs 2 linked genes
This 648 bp open chromatin element is linked to CLDN16 and CLDN1 and is bound by 76 transcription factors.
Linked Genes
2 genes
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
CLDN16 3.4 kb Proximal Proximity
CLDN1 3.5 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr3:190,320,950 – 190,331,598
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
76 transcription factors
Source
Cell type
ATF2 1 dataset
ChIP GM12878 ENCSR000BQK.ATF2.GM12878 148 bp overlap
BATF 1 dataset
ChIP GM12878 ENCFF954REE 231 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 137 bp overlap
BRD4 6 datasets
ChIP CD4_JG1 GSE33281.BRD4.CD4_JG1 272 bp overlap
ChIP HAP1 GSE108387.BRD4.HAP1 648 bp overlap
ChIP RH4_shscr GSE140115.BRD4.RH4_shscr 201 bp overlap
ChIP SUM229PE_pos_30nMtrametinib_24h GSE87418.BRD4.SUM229PE_pos_30nMtrametinib_24h 392 bp overlap
ChIP SUM229PE_pos_DMSO_24h GSE87418.BRD4.SUM229PE_pos_DMSO_24h 551 bp overlap
ChIP keratinocyte GSE140992.BRD4.keratinocyte 648 bp overlap
CHD4 1 dataset
ChIP HaCaT GSE139685.CHD4.HaCaT 169 bp overlap
CREBBP 2 datasets
ChIP keratinocyte GSE140991.CREBBP.keratinocyte 278 bp overlap
ChIP keratinocyte_KLF3i GSE140991.CREBBP.keratinocyte_KLF3i 215 bp overlap
CREM 1 dataset
ChIP GM12878 ENCSR839XZU.CREM.GM12878 175 bp overlap
FEZF1 2 datasets
ChIP HEK293 GSE76494.FEZF1.HEK293 200 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 56 bp overlap
FLI1 1 dataset
ChIP A-673_1_KRAB_GFP GSE106914.FLI1.A-673_1_KRAB_GFP 201 bp overlap
FOS 2 datasets
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 236 bp overlap
ChIP MCF-7 ENCSR569XNP.FOS.MCF-7 224 bp overlap
FOSL2 2 datasets
ChIP NPC_R1159Q GSE122631.FOSL2.NPC_R1159Q 251 bp overlap
ChIP hESC GSE69539.FOSL2.hESC 187 bp overlap
FOXA2 1 dataset
ChIP KerCT GSE90454.FOXA2.KerCT 377 bp overlap
FOXM1 3 datasets
ChIP GM12878 ENCFF264DJE 486 bp overlap
ChIP GM12878 ENCFF264DJE 286 bp overlap
ChIP GM12878 ENCSR000BRU.FOXM1.GM12878 138 bp overlap
GATA2 1 dataset
ChIP ESF GSE108408.GATA2.ESF 86 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 172 bp overlap
JUN 2 datasets
ChIP MCF10A-Er-Src_EtOH GSE115597.JUN.MCF10A-Er-Src_EtOH 200 bp overlap
ChIP keratinocyte_CHD4-KD GSE139685.JUN.keratinocyte_CHD4-KD 151 bp overlap
JUNB 3 datasets
ChIP GM12878 ENCFF667EJQ 227 bp overlap
ChIP GM12878 ENCSR897MMC.JUNB.GM12878 367 bp overlap
ChIP MCF10A-Er-Src_EtOH GSE115597.JUNB.MCF10A-Er-Src_EtOH 164 bp overlap
KDM1A 1 dataset
ChIP NCI-H526_DMSO GSE66297.KDM1A.NCI-H526_DMSO 356 bp overlap
KLF5 2 datasets
ChIP HEK293_E419K GSE88976.KLF5.HEK293_E419K 68 bp overlap
ChIP HEK293_E419Q GSE88976.KLF5.HEK293_E419Q 132 bp overlap
MAML1 1 dataset
ChIP SCC_4h GSE156486.MAML1.SCC_4h 286 bp overlap
MAZ 2 datasets
ChIP HEK293 GSE76494.MAZ.HEK293 193 bp overlap
ChIP HEK293 ENCSR290SSQ.MAZ.HEK293 126 bp overlap
MED1 1 dataset
ChIP hMSC-TERT4_4h GSE104537.MED1.hMSC-TERT4_4h 155 bp overlap
MED12 2 datasets
ChIP leiomyoma_PT886 GSE128230.MED12.leiomyoma_PT886 124 bp overlap
ChIP leiomyoma_PT967 GSE128230.MED12.leiomyoma_PT967 75 bp overlap
MYC 1 dataset
ChIP MCF-10A ENCSR000DOS.MYC.MCF-10A 187 bp overlap
NANOG 3 datasets
ChIP HUES-8 GSE109524.NANOG.HUES-8 89 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 91 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 271 bp overlap
NFIC 1 dataset
ChIP GM12878 ENCSR000BRN.NFIC.GM12878 217 bp overlap
NFKB1 1 dataset
ChIP MCF10A-Er-Src_TAM GSE115597.NFKB1.MCF10A-Er-Src_TAM 180 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 290 bp overlap
NR3C1 3 datasets
ChIP BEAS-2B_TNF_IA1 GSE125623.NR3C1.BEAS-2B_TNF_IA1 491 bp overlap
ChIP HCC1937 GSE152203.NR3C1.HCC1937 150 bp overlap
ChIP SUM159PT GSE152203.NR3C1.SUM159PT 156 bp overlap
PAX7 1 dataset
ChIP H9_DOX GSE98976.PAX7.H9_DOX 213 bp overlap
PGR 1 dataset
ChIP T-47D_R5020-MTVL GSE126859.PGR.T-47D_R5020-MTVL 168 bp overlap
PML 1 dataset
ChIP GM12878 ENCSR000BQM.PML.GM12878 142 bp overlap
POLR2A 3 datasets
ChIP esophagus squamous epithelium ENCFF708IOX 284 bp overlap
ChIP vagina ENCFF305NWS 439 bp overlap
ChIP vagina ENCFF305NWS 477 bp overlap
POU2F2 1 dataset
ChIP GM12878 ENCFF207RKY 263 bp overlap
POU2F3 1 dataset
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 441 bp overlap
POU5F1 2 datasets
ChIP HUES-8 GSE109524.POU5F1.HUES-8 107 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 91 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 327 bp overlap
RBPJ 1 dataset
ChIP SCC_4h GSE156486.RBPJ.SCC_4h 266 bp overlap
RELA 8 datasets
ChIP Detroit-562_LPS GSE91018.RELA.Detroit-562_LPS 367 bp overlap
ChIP Detroit-562_Pam2CSK4 GSE91018.RELA.Detroit-562_Pam2CSK4 296 bp overlap
ChIP Detroit-562_Poly-I GSE91018.RELA.Detroit-562_Poly-I 354 bp overlap
ChIP Detroit-562_TNFa GSE91018.RELA.Detroit-562_TNFa 377 bp overlap
ChIP Detroit-562_tri-DAP GSE91018.RELA.Detroit-562_tri-DAP 355 bp overlap
ChIP FaDu_LPS GSE132018.RELA.FaDu_LPS 648 bp overlap
ChIP HDF_NUTLIN GSE77225.RELA.HDF_NUTLIN 289 bp overlap
ChIP aortic-endothelial-cell_IL1B_D2 GSE139377.RELA.aortic-endothelial-cell_IL1B_D2 201 bp overlap
RUNX3 1 dataset
ChIP GM12878 ENCFF395WHA 326 bp overlap
SIX2 3 datasets
ChIP HEK GSE73865.SIX2.HEK 75 bp overlap
ChIP kidney_fetal_16w GSE75948.SIX2.kidney_fetal_16w 57 bp overlap
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 73 bp overlap
SMAD3 2 datasets
ChIP HCC1954 GSE104760.SMAD3.HCC1954 519 bp overlap
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 238 bp overlap
SMARCA4 5 datasets
ChIP A-549_AG15686 GSE132290.SMARCA4.A-549_AG15686 237 bp overlap
ChIP A-549_AG15688 GSE132290.SMARCA4.A-549_AG15688 283 bp overlap
ChIP A-549_AG15725 GSE132290.SMARCA4.A-549_AG15725 57 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 534 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 159 bp overlap
SOX2 1 dataset
ChIP KYSE-70 GSE46837.SOX2.KYSE-70 287 bp overlap
SP4 1 dataset
ChIP HEK293 GSE76494.SP4.HEK293 204 bp overlap
TBP 1 dataset
ChIP GM12878 ENCFF571OXR 385 bp overlap
TBX21 1 dataset
ChIP GM12878 ENCSR739IHN.TBX21.GM12878 260 bp overlap
TEAD1 1 dataset
ChIP keratinocyte GSE138727.TEAD1.keratinocyte 293 bp overlap
TFAP2C 2 datasets
ChIP HCC1954 GSE126897.TFAP2C.HCC1954 186 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 266 bp overlap
TP53 44 datasets
ChIP A-498_2h_4GY GSE100292.TP53.A-498_2h_4GY 203 bp overlap
ChIP GM00011 GSE55727.TP53.GM00011 648 bp overlap
ChIP GM06170 GSE55727.TP53.GM06170 648 bp overlap
ChIP H9 GSE142050.TP53.H9 492 bp overlap
ChIP H9_ectoderm GSE142050.TP53.H9_ectoderm 383 bp overlap
ChIP H9_mesoderm GSE142050.TP53.H9_mesoderm 483 bp overlap
ChIP HCT-116_5FU GSE58506.TP53.HCT-116_5FU 278 bp overlap
ChIP HCT-116_Nutlin3a-SC GSE125927.TP53.HCT-116_Nutlin3a-SC 200 bp overlap
ChIP HCT-116_nutlin GSE86164.TP53.HCT-116_nutlin 563 bp overlap
ChIP IMR-90 GSE115940.TP53.IMR-90 648 bp overlap
ChIP IMR-90_APO GSE53491.TP53.IMR-90_APO 188 bp overlap
ChIP IMR-90_DMSO GSE58740.TP53.IMR-90_DMSO 155 bp overlap
ChIP IMR-90_NUT3A GSE58740.TP53.IMR-90_NUT3A 648 bp overlap
ChIP IMR-90_SENES_SHLUC GSE42728.TP53.IMR-90_SENES_SHLUC 146 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP53.MCF-10A_Nutlin3A 590 bp overlap
ChIP MCF-7_1h_IR_10Gy GSE100099.TP53.MCF-7_1h_IR_10Gy 321 bp overlap
ChIP MCF-7_2-5h_IR_10Gy GSE100099.TP53.MCF-7_2-5h_IR_10Gy 347 bp overlap
ChIP MCF-7_7-5h_IR_10Gy GSE100099.TP53.MCF-7_7-5h_IR_10Gy 487 bp overlap
ChIP MCF-7_7-5h_IR_10Gy_Nutlin GSE100099.TP53.MCF-7_7-5h_IR_10Gy_Nutlin 540 bp overlap
ChIP MCF-7_NCS-treated GSE101737.TP53.MCF-7_NCS-treated 237 bp overlap
ChIP MCF-7_nutlin GSE86164.TP53.MCF-7_nutlin 648 bp overlap
ChIP MOLM-13_R282W_DMSO GSE131484.TP53.MOLM-13_R282W_DMSO 432 bp overlap
ChIP MOLM-13_R282W_Daunorubicin GSE131484.TP53.MOLM-13_R282W_Daunorubicin 377 bp overlap
ChIP SJSA-1 GSE86164.TP53.SJSA-1 482 bp overlap
ChIP SJSA-1_nutlin GSE86164.TP53.SJSA-1_nutlin 648 bp overlap
ChIP SaOS-2 GSE15780.TP53.SaOS-2 592 bp overlap
ChIP U2OS_DMSO GSE46641.TP53.U2OS_DMSO 172 bp overlap
ChIP U2OS_DXR GSE46641.TP53.U2OS_DXR 238 bp overlap
ChIP U2OS_NUT GSE46641.TP53.U2OS_NUT 475 bp overlap
ChIP U2OS_UV_16H ERP004176.TP53.U2OS_UV_16H 308 bp overlap
ChIP U2OS_UV_8H ERP004176.TP53.U2OS_UV_8H 212 bp overlap
ChIP UACC-257_nutlin_2h GSE100292.TP53.UACC-257_nutlin_2h 289 bp overlap
ChIP UO-31_2h_4GY GSE100292.TP53.UO-31_2h_4GY 595 bp overlap
ChIP keratinocyte GSE56674.TP53.keratinocyte 254 bp overlap
ChIP keratinocyte_ADRIA GSE56674.TP53.keratinocyte_ADRIA 352 bp overlap
ChIP keratinocyte_CISP GSE56674.TP53.keratinocyte_CISP 428 bp overlap
ChIP lymphocyte_104_Nutlin GSE110368.TP53.lymphocyte_104_Nutlin 390 bp overlap
ChIP lymphocyte_116 GSE110368.TP53.lymphocyte_116 260 bp overlap
ChIP lymphocyte_116_DXR GSE110368.TP53.lymphocyte_116_DXR 480 bp overlap
ChIP lymphocyte_116_Nutlin GSE110368.TP53.lymphocyte_116_Nutlin 627 bp overlap
ChIP lymphocyte_45_DXR GSE110368.TP53.lymphocyte_45_DXR 420 bp overlap
ChIP lymphocyte_45_Nutlin GSE110368.TP53.lymphocyte_45_Nutlin 246 bp overlap
ChIP lymphocyte_90_DXR GSE110368.TP53.lymphocyte_90_DXR 588 bp overlap
ChIP lymphocyte_90_Nutlin GSE110368.TP53.lymphocyte_90_Nutlin 506 bp overlap
TP53_PS15 2 datasets
ChIP U2OS_ACTD GSE21939.TP53_PS15.U2OS_ACTD 295 bp overlap
ChIP U2OS_ETO GSE21939.TP53_PS15.U2OS_ETO 329 bp overlap
TP63 22 datasets
ChIP BxPC-3 GSE115461.TP63.BxPC-3 576 bp overlap
ChIP EP156T GSE43111.TP63.EP156T 239 bp overlap
ChIP HCC95 GSE46837.TP63.HCC95 244 bp overlap
ChIP HaCaT_LacZ GSE60814.TP63.HaCaT_LacZ 195 bp overlap
ChIP HaCaT_LacZ_TGFB GSE60814.TP63.HaCaT_LacZ_TGFB 486 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 484 bp overlap
ChIP HaCaT_caRAS_TGFB GSE60814.TP63.HaCaT_caRAS_TGFB 122 bp overlap
ChIP HaCaT_dnRAS_TGFB GSE60814.TP63.HaCaT_dnRAS_TGFB 492 bp overlap
ChIP JHU-029 GSE88859.TP63.JHU-029 365 bp overlap
ChIP KYSE-70 GSE46837.TP63.KYSE-70 368 bp overlap
ChIP MCF-10A_DCIS GSE72009.TP63.MCF-10A_DCIS 383 bp overlap
ChIP MCF-10A_Nutlin3A GSE111009.TP63.MCF-10A_Nutlin3A 648 bp overlap
ChIP SUIT-2 GSE115461.TP63.SUIT-2 648 bp overlap
ChIP keratinocyte GSE56674.TP63.keratinocyte 389 bp overlap
ChIP keratinocyte_ADRIA GSE56674.TP63.keratinocyte_ADRIA 263 bp overlap
ChIP keratinocyte_CISP GSE56674.TP63.keratinocyte_CISP 369 bp overlap
ChIP keratinocyte_D0 GSE59824.TP63.keratinocyte_D0 441 bp overlap
ChIP keratinocyte_D2 GSE59824.TP63.keratinocyte_D2 415 bp overlap
ChIP keratinocyte_D4 GSE59824.TP63.keratinocyte_D4 358 bp overlap
ChIP keratinocyte_D7 GSE59824.TP63.keratinocyte_D7 370 bp overlap
ChIP keratinocyte_epidermal GSE67382.TP63.keratinocyte_epidermal 523 bp overlap
ChIP keratinocyte_epidermal_KDPAF GSE67382.TP63.keratinocyte_epidermal_KDPAF 304 bp overlap
TP73 1 dataset
ChIP GM12878 GSE97661.TP73.GM12878 293 bp overlap
TP73_TA 1 dataset
ChIP SaOS-2 GSE15780.TP73_TA.SaOS-2 315 bp overlap
TRIM28 1 dataset
ChIP HEK293 ENCFF582MWI 568 bp overlap
WT1 2 datasets
ChIP HEK293 ENCFF906HIR 65 bp overlap
ChIP HEK293 ENCSR966PJJ.WT1.HEK293 161 bp overlap
YY1 3 datasets
ChIP HEK293 ENCSR859RAO.YY1.HEK293 276 bp overlap
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 142 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 498 bp overlap
ZBED1 2 datasets
ChIP GM12878 ENCFF007OSW 440 bp overlap
ChIP GM12878 ENCFF007OSW 505 bp overlap
ZBTB17 1 dataset
ChIP HEK293 ENCFF865LIO 441 bp overlap
ZFP37 3 datasets
ChIP HEK293 ENCFF968PWB 491 bp overlap
ChIP HEK293 ENCFF968PWB 366 bp overlap
ChIP HEK293 ENCSR365GRX.ZFP37.HEK293 314 bp overlap
ZNF335 2 datasets
ChIP HEK293 ENCFF784SLD 577 bp overlap
ChIP HEK293 ENCSR328SUD.ZNF335.HEK293 448 bp overlap
ZNF341 1 dataset
ChIP HEK293 GSE76494.ZNF341.HEK293 81 bp overlap
ZNF823 1 dataset
ChIP HEK293T GSE78099.ZNF823.HEK293T 290 bp overlap