chr4 : 56,055,093 – 56,055,833
740 bp 115 TFs 3 linked genes
This 740 bp open chromatin element is linked to CRACD, CEP135, and EXOC1 and is bound by 115 transcription factors.
Linked Genes
3 genes
Distance
Link type
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
CRACD 6.0 kb Proximal Proximity
CEP135 106.5 kb Distal Multiome
EXOC1 201.7 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:56,050,093 – 56,060,833
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
115 transcription factors
Source
Cell type
AR 1 dataset
ChIP LNCaP_1F5_SIFOXA1 GSE30623.AR.LNCaP_1F5_SIFOXA1 143 bp overlap
ARID3A 1 dataset
ChIP HepG2 ENCFF341DES 272 bp overlap
CEBPA 2 datasets
ChIP Kasumi-1_CEBPA-ER GSE102697.CEBPA.Kasumi-1_CEBPA-ER 158 bp overlap
ChIP Kasumi-1_CEBPA-ER_E2 GSE102697.CEBPA.Kasumi-1_CEBPA-ER_E2 343 bp overlap
CHD4 1 dataset
ChIP SCMC GSE155861.CHD4.SCMC 160 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 210 bp overlap
Cebpa 4 datasets
ChIP BLaER1 ENCFF093OYK 437 bp overlap
ChIP BLaER1 ENCFF335XTP 257 bp overlap
ChIP BLaER1 ENCFF346MCV 136 bp overlap
ChIP BLaER1 ENCFF460KDD 397 bp overlap
E2F6 3 datasets
Motif DE_36h DE_36h-E2F6_MA0471.3 8 bp overlap
Motif DE_48h DE_48h-E2F6_MA0471.3 8 bp overlap
Motif DE_60h DE_60h-E2F6_MA0471.3 8 bp overlap
EBF1 2 datasets
Motif DE_36h DE_36h-EBF1_MA0154.5 11 bp overlap
Motif DE_60h DE_60h-EBF1_MA0154.5 11 bp overlap
EGR1 1 dataset
ChIP fibroblast_DOX-24h GSE134924.EGR1.fibroblast_DOX-24h 200 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 463 bp overlap
ETS1 1 dataset
ChIP CTV-1_DSG GSE128835.ETS1.CTV-1_DSG 172 bp overlap
EWSR1-FLI1 4 datasets
Motif DE_36h DE_36h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_48h DE_48h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_60h DE_60h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_72h DE_72h-EWSR1-FLI1_MA0149.1 18 bp overlap
Ebf4 2 datasets
Motif DE_36h DE_36h-Ebf4_MA2122.1 11 bp overlap
Motif DE_60h DE_60h-Ebf4_MA2122.1 11 bp overlap
Elf5 4 datasets
Motif DE_36h DE_36h-Elf5_MA0136.4 8 bp overlap
Motif DE_48h DE_48h-Elf5_MA0136.4 8 bp overlap
Motif DE_60h DE_60h-Elf5_MA0136.4 8 bp overlap
Motif DE_72h DE_72h-Elf5_MA0136.4 8 bp overlap
FIGLA 2 datasets
Motif DE_36h DE_36h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 239 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 704 bp overlap
ChIP DE DE-FOXA2-2 607 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 339 bp overlap
GATA1 5 datasets
ChIP K-562_dCas9-LSD1 GSE132212.GATA1.K-562_dCas9-LSD1 145 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 197 bp overlap
ChIP erythroblast ENCFF867JAR 522 bp overlap
ChIP erythroblast ENCSR000EXP.GATA1.erythroblast 350 bp overlap
ChIP erythroid-progenitor GSE124163.GATA1.erythroid-progenitor 156 bp overlap
GATA2 6 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 166 bp overlap
GATA3 1 dataset
ChIP MCF-7_E2 GSE81510.GATA3.MCF-7_E2 210 bp overlap
GATA4 10 datasets
ChIP DE DE-GATA4-1 735 bp overlap
ChIP DE DE-GATA4-2 720 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 345 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 532 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 740 bp overlap
GATA5 5 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 22 datasets
ChIP AGS GSE51705.GATA6.AGS 193 bp overlap
ChIP DE DE-GATA6-1 620 bp overlap
ChIP DE DE-GATA6-2 740 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 489 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 432 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 640 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 503 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 620 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 673 bp overlap
ChIP HUG1N GSE51936.GATA6.HUG1N 164 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 226 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 466 bp overlap
ChIP foregut GSE117136.GATA6.foregut 392 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 397 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 373 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 289 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 505 bp overlap
GRHL1 1 dataset
Motif DE_36h DE_36h-GRHL1_MA0647.2 10 bp overlap
GRHL2 1 dataset
Motif DE_36h DE_36h-GRHL2_MA1105.3 8 bp overlap
Gata3 5 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HAND2 1 dataset
ChIP SK-N-BE2-C GSE94822.HAND2.SK-N-BE2-C 109 bp overlap
HNF1A 1 dataset
Motif DE_36h DE_36h-HNF1A_MA0046.3 13 bp overlap
HNF1B 1 dataset
Motif DE_36h DE_36h-HNF1B_MA0153.2 13 bp overlap
HNF4A 4 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 104 bp overlap
ChIP Hep-G2 ENCSR000BLF.HNF4A.Hep-G2 173 bp overlap
ChIP Hep-G2 ENCSR469FBY.HNF4A.Hep-G2 159 bp overlap
ChIP gastric-epithelial-cell_OE GSE114018.HNF4A.gastric-epithelial-cell_OE 69 bp overlap
HOXB13 1 dataset
ChIP G-401 GSE65381.HOXB13.G-401 187 bp overlap
Hmga1 1 dataset
Motif DE_36h DE_36h-Hmga1_MA2124.1 8 bp overlap
IKZF2 4 datasets
Motif DE_36h DE_36h-IKZF2_MA2326.1 6 bp overlap
Motif DE_48h DE_48h-IKZF2_MA2326.1 6 bp overlap
Motif DE_60h DE_60h-IKZF2_MA2326.1 6 bp overlap
Motif DE_72h DE_72h-IKZF2_MA2326.1 6 bp overlap
KDM1A 1 dataset
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 137 bp overlap
KMT2A 4 datasets
ChIP HEK293T_N-term_C104 GSE90762.KMT2A.HEK293T_N-term_C104 404 bp overlap
ChIP HEK293T_N-term_C49 GSE90762.KMT2A.HEK293T_N-term_C49 313 bp overlap
ChIP HEK293T_N-term_shCtrl_TASP1-KO GSE90762.KMT2A.HEK293T_N-term_shCtrl_TASP1-KO 441 bp overlap
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 175 bp overlap
LIN54 1 dataset
Motif DE_36h DE_36h-LIN54_MA0619.2 7 bp overlap
Lhx3 1 dataset
Motif DE_36h DE_36h-Lhx3_MA0135.2 12 bp overlap
MAZ 1 dataset
ChIP HEK293 GSE76494.MAZ.HEK293 210 bp overlap
MEF2B 1 dataset
ChIP tonsil GSE110682.MEF2B.tonsil 58 bp overlap
MIXL1 1 dataset
ChIP HepG2 ENCFF817YFO 313 bp overlap
MYC 1 dataset
ChIP Jurkat GSE83777.MYC.Jurkat 192 bp overlap
Mecom 1 dataset
Motif DE_36h DE_36h-Mecom_MA0029.2 11 bp overlap
NEUROD1 4 datasets
ChIP D283-Med GSE92582.NEUROD1.D283-Med 293 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 185 bp overlap
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 106 bp overlap
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 168 bp overlap
NFYB 1 dataset
Motif DE_36h DE_36h-NFYB_MA0502.3 9 bp overlap
NKX2-2 1 dataset
Motif DE_36h DE_36h-NKX2-2_MA1645.2 8 bp overlap
NOTCH1 1 dataset
ChIP GSC8-11_dasatinib GSE74557.NOTCH1.GSC8-11_dasatinib 63 bp overlap
NR2F2 1 dataset
ChIP Hep-G2 ENCSR000BVM.NR2F2.Hep-G2 125 bp overlap
NR2F6 2 datasets
Motif DE_36h DE_36h-NR2F6_MA1539.1 15 bp overlap
ChIP Hep-G2 ENCSR518WPL.NR2F6.Hep-G2 175 bp overlap
ONECUT1 3 datasets
ChIP H9 ERP004206.ONECUT1.H9 142 bp overlap
ChIP Hep-G2 ENCSR956OSX.ONECUT1.Hep-G2 107 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 119 bp overlap
PATZ1 1 dataset
ChIP HEK293 ENCSR966ULI.PATZ1.HEK293 399 bp overlap
PDX1 3 datasets
ChIP H9 ERP004206.PDX1.H9 163 bp overlap
ChIP hiPSC GSE125768.PDX1.hiPSC 309 bp overlap
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 192 bp overlap
PHIP 1 dataset
ChIP HEK293_ab833 GSE101646.PHIP.HEK293_ab833 292 bp overlap
PHOX2A 1 dataset
Motif DE_36h DE_36h-PHOX2A_MA0713.1 11 bp overlap
PHOX2B 1 dataset
Motif DE_36h DE_36h-PHOX2B_MA0681.3 12 bp overlap
PITX3 1 dataset
ChIP SH-SY5Y GSE93275.PITX3.SH-SY5Y 191 bp overlap
POU2F1 1 dataset
Motif DE_36h DE_36h-POU2F1_MA0785.2 9 bp overlap
POU3F1 1 dataset
Motif DE_36h DE_36h-POU3F1_MA0786.2 10 bp overlap
POU3F2 1 dataset
Motif DE_36h DE_36h-POU3F2_MA0787.1 12 bp overlap
POU3F3 1 dataset
Motif DE_36h DE_36h-POU3F3_MA0788.1 13 bp overlap
POU4F2 1 dataset
ChIP HNPC_DIF GSE74814.POU4F2.HNPC_DIF 92 bp overlap
PRDM10 1 dataset
ChIP HEK293 ENCFF145WQQ 263 bp overlap
PRDM15 1 dataset
ChIP WTC11 ENCFF108TMF 342 bp overlap
PRDM6 1 dataset
ChIP HEK293 ENCFF283AJL 107 bp overlap
PROP1 1 dataset
Motif DE_36h DE_36h-PROP1_MA0715.1 11 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 210 bp overlap
Plagl1 1 dataset
Motif DE_36h DE_36h-Plagl1_MA1615.2 8 bp overlap
Prdm4 1 dataset
Motif DE_36h DE_36h-Prdm4_MA1647.3 11 bp overlap
RARA 1 dataset
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 248 bp overlap
RBPJ 1 dataset
ChIP GSC8-11_dasatinib GSE74557.RBPJ.GSC8-11_dasatinib 69 bp overlap
RELA 1 dataset
ChIP SGBS_TNF GSE64233.RELA.SGBS_TNF 467 bp overlap
SCRT2 1 dataset
Motif DE_36h DE_36h-SCRT2_MA0744.3 10 bp overlap
SIX2 1 dataset
ChIP HEK GSE73865.SIX2.HEK 170 bp overlap
SMAD2 1 dataset
ChIP endoderm GSE29422.SMAD2.endoderm 148 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 566 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 405 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 325 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 511 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 552 bp overlap
SMAD3 1 dataset
ChIP endoderm GSE29422.SMAD3.endoderm 225 bp overlap
SMAD4 1 dataset
ChIP endoderm GSE29422.SMAD4.endoderm 222 bp overlap
SMARCA4 9 datasets
ChIP CTV-1_DSG GSE128835.SMARCA4.CTV-1_DSG 293 bp overlap
ChIP CTV-1_FLAG GSE128835.SMARCA4.CTV-1_FLAG 244 bp overlap
ChIP CTV-1_delQ GSE128835.SMARCA4.CTV-1_delQ 182 bp overlap
ChIP NPC_K755R-siAP GSE122631.SMARCA4.NPC_K755R-siAP 198 bp overlap
ChIP NPC_K755R-siCT GSE122631.SMARCA4.NPC_K755R-siCT 328 bp overlap
ChIP NPC_siCT GSE122631.SMARCA4.NPC_siCT 334 bp overlap
ChIP NPC_siSOX GSE122631.SMARCA4.NPC_siSOX 253 bp overlap
ChIP NPC_siSOX GSE122631.SMARCA4.NPC_siSOX 183 bp overlap
ChIP NSC GSE125033.SMARCA4.NSC 451 bp overlap
SMARCB1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 170 bp overlap
SMARCC1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 212 bp overlap
SOX13 1 dataset
ChIP HepG2 ENCFF062VSQ 267 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 454 bp overlap
SOX2 1 dataset
ChIP HNSC GSE69479.SOX2.HNSC 185 bp overlap
SP1 2 datasets
Motif DE_36h DE_36h-SP1_MA0079.5 9 bp overlap
Motif DE_60h DE_60h-SP1_MA0079.5 9 bp overlap
SP2 3 datasets
Motif DE_36h DE_36h-SP2_MA0516.3 9 bp overlap
Motif DE_60h DE_60h-SP2_MA0516.3 9 bp overlap
ChIP K-562 ENCSR000BNL.SP2.K-562 113 bp overlap
SP5 2 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
SREBF1 1 dataset
Motif DE_36h DE_36h-SREBF1_MA0595.1 10 bp overlap
SREBF2 1 dataset
Motif DE_36h DE_36h-SREBF2_MA0596.1 10 bp overlap
SRF 2 datasets
ChIP Hep-G2 ENCSR000BLV.SRF.Hep-G2 156 bp overlap
ChIP HepG2 ENCFF625QHW 185 bp overlap
STAT3 1 dataset
ChIP MCF-7_jc5836 GSE126004.STAT3.MCF-7_jc5836 251 bp overlap
SUZ12 1 dataset
ChIP CRL-7250_shSS18 GSE108025.SUZ12.CRL-7250_shSS18 296 bp overlap
TFDP1 3 datasets
Motif DE_36h DE_36h-TFDP1_MA1122.2 8 bp overlap
Motif DE_48h DE_48h-TFDP1_MA1122.2 8 bp overlap
Motif DE_60h DE_60h-TFDP1_MA1122.2 8 bp overlap
THRB 1 dataset
ChIP Hep-G2 ENCSR430JGJ.THRB.Hep-G2 190 bp overlap
TRPS1 5 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
TWIST1 2 datasets
ChIP BE2C GSE80151.TWIST1.BE2C 113 bp overlap
ChIP SK-N-BE2-C GSE80151.TWIST1.SK-N-BE2-C 113 bp overlap
YY1 2 datasets
ChIP Hep-G2_NC GSE120104.YY1.Hep-G2_NC 314 bp overlap
ChIP Hep-G2_RBM25-KD GSE120104.YY1.Hep-G2_RBM25-KD 489 bp overlap
ZBTB26 1 dataset
Motif DE_36h DE_36h-ZBTB26_MA1579.2 8 bp overlap
ZEB1 3 datasets
Motif DE_36h DE_36h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR418KUS.ZEB1.neuron_bipolar_doxy_4d 193 bp overlap
ZNF207 1 dataset
ChIP WA09 GSE118632.ZNF207.WA09 245 bp overlap
ZNF467 1 dataset
ChIP HEK293 GSE76494.ZNF467.HEK293 148 bp overlap
ZNF652 2 datasets
ChIP Hep-G2 ENCSR502GAX.ZNF652.Hep-G2 665 bp overlap
ChIP HepG2 ENCFF331VPZ 195 bp overlap
ZNF770 2 datasets
ChIP HEK293 ENCSR242BGR.ZNF770.HEK293 302 bp overlap
ChIP HEK293 GSE76494.ZNF770.HEK293 203 bp overlap