chr4 : 56,787,104 – 56,787,870
766 bp 101 TFs 3 linked genes
This 766 bp open chromatin element is linked to REST, NOA1, and POLR2B and is bound by 101 transcription factors.
Linked Genes
3 genes
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
REST 120.4 kb Distal Multiome
NOA1 190.2 kb Distal Multiome
POLR2B 190.3 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:56,782,104 – 56,792,870
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
101 transcription factors
Source
Cell type
AR 4 datasets
ChIP fibroblast_prostate-cancer_PCDF2 GSE126852.AR.fibroblast_prostate-cancer_PCDF2 221 bp overlap
ChIP prostate GSE56288.AR.prostate 256 bp overlap
ChIP prostate_cancer-associated-fibroblasts GSE90772.AR.prostate_cancer-associated-fibroblasts 289 bp overlap
ChIP prostate_fetal-fibroblast GSE90772.AR.prostate_fetal-fibroblast 225 bp overlap
ASCL1 1 dataset
ChIP NCI-H82 GSE69394.ASCL1.NCI-H82 162 bp overlap
BCOR 1 dataset
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 324 bp overlap
BRD2 1 dataset
ChIP SUM159PT_MGSK2801 GSE116879.BRD2.SUM159PT_MGSK2801 232 bp overlap
BRD4 13 datasets
ChIP HEK293T GSE51633.BRD4.HEK293T 135 bp overlap
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 84 bp overlap
ChIP K-562_DMSO GSE99178.BRD4.K-562_DMSO 190 bp overlap
ChIP Kelly_resistant_JQ1 GSE103030.BRD4.Kelly_resistant_JQ1 224 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 502 bp overlap
ChIP MDA-MB-231_JQ1_MGSK2801 GSE116879.BRD4.MDA-MB-231_JQ1_MGSK2801 232 bp overlap
ChIP MDA-MB-231_JQ1_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_JQ1_MGSK2801_rDNA 232 bp overlap
ChIP MPNST_PDJQ GSE62499.BRD4.MPNST_PDJQ 112 bp overlap
ChIP SUM159PT_DMSO_24h GSE87418.BRD4.SUM159PT_DMSO_24h 228 bp overlap
ChIP SUM159PT_DMSO_8h GSE87418.BRD4.SUM159PT_DMSO_8h 284 bp overlap
ChIP SUM159PT_Dox_48h GSE87418.BRD4.SUM159PT_Dox_48h 137 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 598 bp overlap
ChIP retina_AB1-RB GSE86981.BRD4.retina_AB1-RB 244 bp overlap
CBFA2T3 2 datasets
ChIP K-562 ENCSR697YLJ.CBFA2T3.K-562 345 bp overlap
ChIP K562 ENCFF673OEZ 411 bp overlap
CDK9 1 dataset
ChIP HEK293T_SICTR GSE51633.CDK9.HEK293T_SICTR 155 bp overlap
CDX2 4 datasets
ChIP Caco-2_DIFF GSE23436.CDX2.Caco-2_DIFF 139 bp overlap
ChIP LS180 GSE31939.CDX2.LS180 210 bp overlap
ChIP LS180_125 GSE31939.CDX2.LS180_125 249 bp overlap
ChIP adult-duodenal-cell GSE115314.CDX2.adult-duodenal-cell 244 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_neural-crest-cell GSE108506.CHD7.hiPSC_derived_neural-crest-cell 176 bp overlap
CTBP1 1 dataset
ChIP K-562 ENCSR201NQZ.CTBP1.K-562 490 bp overlap
CTCF 3 datasets
ChIP Kasumi-1_ctrl GSE121280.CTCF.Kasumi-1_ctrl 123 bp overlap
ChIP THP-1_macrophage_PMA GSE96800.CTCF.THP-1_macrophage_PMA 220 bp overlap
ChIP anterior-temporal-cortex_adult GSE116825.CTCF.anterior-temporal-cortex_adult 353 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF262VBH 251 bp overlap
E2F1 1 dataset
ChIP U-87MG_GBM GSE99171.E2F1.U-87MG_GBM 210 bp overlap
E2F7 1 dataset
ChIP IMR-90_SENES_SHCTR GSE40343.E2F7.IMR-90_SENES_SHCTR 52 bp overlap
EGR1 1 dataset
ChIP T-HESCs GSE141063.EGR1.T-HESCs 226 bp overlap
ELK1 1 dataset
ChIP K-562 ENCSR338QAC.ELK1.K-562 199 bp overlap
EP300 1 dataset
ChIP SK-N-SH ENCSR000BUA.EP300.SK-N-SH 57 bp overlap
ERG 7 datasets
ChIP K-562_DOX GSE23730.ERG.K-562_DOX 270 bp overlap
ChIP SKNO-1 GSE23730.ERG.SKNO-1 402 bp overlap
ChIP VCaP GSE83650.ERG.VCaP 168 bp overlap
ChIP VCaP GSE98809.ERG.VCaP 168 bp overlap
ChIP VCaP_DHAT_18H GSE28950.ERG.VCaP_DHAT_18H 135 bp overlap
ChIP VCaP_DHT GSE79128.ERG.VCaP_DHT 420 bp overlap
ChIP VCaP_shARID1A GSE110655.ERG.VCaP_shARID1A 153 bp overlap
ESRRG 1 dataset
ChIP cardiomyocyte GSE113760.ESRRG.cardiomyocyte 205 bp overlap
ETV1 4 datasets
ChIP GIST-T1 GSE106624.ETV1.GIST-T1 312 bp overlap
ChIP GIST-T1 GSE80443.ETV1.GIST-T1 184 bp overlap
ChIP GIST-T1 GSE106624.ETV1.GIST-T1 365 bp overlap
ChIP GIST-T1 GSE80443.ETV1.GIST-T1 168 bp overlap
FEZF1 2 datasets
ChIP HEK293 ENCFF528YED 376 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 302 bp overlap
FLI1 1 dataset
ChIP SKNO-1 GSE23730.FLI1.SKNO-1 533 bp overlap
FOS 3 datasets
ChIP IMR-90 ENCFF179EDA 128 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 79 bp overlap
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 67 bp overlap
FOSL2 4 datasets
ChIP NPC_K755R-pos GSE122631.FOSL2.NPC_K755R-pos 111 bp overlap
ChIP SK-N-SH ENCFF127ZDW 95 bp overlap
ChIP SK-N-SH ENCSR000BVB.FOSL2.SK-N-SH 58 bp overlap
ChIP hESC GSE69539.FOSL2.hESC 75 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 327 bp overlap
ChIP DE DE-FOXA2-2 294 bp overlap
FOXF1 2 datasets
ChIP GIST-T1 GSE106624.FOXF1.GIST-T1 185 bp overlap
ChIP GIST-T1 GSE106624.FOXF1.GIST-T1 147 bp overlap
FOXL2 2 datasets
ChIP HGrC1_C134W-TGF GSE138496.FOXL2.HGrC1_C134W-TGF 620 bp overlap
ChIP HGrC1_C134W-TGF_parental GSE138496.FOXL2.HGrC1_C134W-TGF_parental 274 bp overlap
GABPA 1 dataset
ChIP K562 ENCFF139LXS 674 bp overlap
GATA2 6 datasets
ChIP ME-1_CBFB-MYH11-KD GSE117138.GATA2.ME-1_CBFB-MYH11-KD 283 bp overlap
ChIP SKH1 GSE87283.GATA2.SKH1 217 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 567 bp overlap
ChIP dermal-fibroblast_alone GSE51025.GATA2.dermal-fibroblast_alone 122 bp overlap
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 396 bp overlap
ChIP hiPSC_SLC9ebs GSE107639.GATA2.hiPSC_SLC9ebs 189 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 524 bp overlap
ChIP DE DE-GATA4-2 582 bp overlap
ChIP foregut GSE117136.GATA4.foregut 317 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 530 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 604 bp overlap
GATA6 14 datasets
ChIP DE DE-GATA6-1 614 bp overlap
ChIP DE DE-GATA6-2 599 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 514 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 452 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 518 bp overlap
ChIP ESO-26 GSE132680.GATA6.ESO-26 723 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 500 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 588 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 515 bp overlap
ChIP foregut GSE117136.GATA6.foregut 414 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 352 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 265 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 295 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 271 bp overlap
GLIS1 2 datasets
ChIP HEK293 ENCFF299RSE 268 bp overlap
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 532 bp overlap
HAND2 1 dataset
ChIP Kelly GSE94822.HAND2.Kelly 235 bp overlap
HDAC1 1 dataset
ChIP K-562 GSE140325.HDAC1.K-562 229 bp overlap
HDAC2 2 datasets
ChIP K-562 ENCSR000BMG.HDAC2.K-562 197 bp overlap
ChIP RH4_Entinostat-6H GSE116344.HDAC2.RH4_Entinostat-6H 318 bp overlap
HDAC3 1 dataset
ChIP RH4_DMSO-6H GSE116344.HDAC3.RH4_DMSO-6H 414 bp overlap
HNF4A 1 dataset
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 116 bp overlap
IKZF1 3 datasets
ChIP K-562 ENCSR395HWC.IKZF1.K-562 639 bp overlap
ChIP K562 ENCFF348IBL 456 bp overlap
ChIP K562 ENCFF771OHZ 497 bp overlap
IKZF3 1 dataset
ChIP HEK293 ENCSR304AMN.IKZF3.HEK293 254 bp overlap
KLF5 1 dataset
ChIP ESO-26 GSE132680.KLF5.ESO-26 285 bp overlap
MAZ 2 datasets
ChIP HEK293 GSE76494.MAZ.HEK293 91 bp overlap
ChIP IMR-90 ENCSR000EFF.MAZ.IMR-90 209 bp overlap
MED1 1 dataset
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 264 bp overlap
MED12 2 datasets
ChIP leiomyoma_PT886 GSE128230.MED12.leiomyoma_PT886 128 bp overlap
ChIP leiomyoma_PT967 GSE128230.MED12.leiomyoma_PT967 58 bp overlap
MYC 1 dataset
ChIP GP5D GSE51234.MYC.GP5D 445 bp overlap
MYCN 1 dataset
ChIP Kelly GSE94822.MYCN.Kelly 214 bp overlap
MYOD1 4 datasets
ChIP IMR-90_GM GSE128527.MYOD1.IMR-90_GM 382 bp overlap
ChIP RH4 GSE83726.MYOD1.RH4 297 bp overlap
ChIP myoblast GSE50413.MYOD1.myoblast 144 bp overlap
ChIP rhabdomyosarcoma GSE50413.MYOD1.rhabdomyosarcoma 65 bp overlap
NANOG 1 dataset
ChIP HUES-8 GSE109524.NANOG.HUES-8 303 bp overlap
NCOA2 1 dataset
ChIP MCF-7 ERP000901.NCOA2.MCF-7 163 bp overlap
NCOR1 1 dataset
ChIP K-562 ENCSR798ILC.NCOR1.K-562 285 bp overlap
NEUROD1 1 dataset
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 550 bp overlap
NEUROG2 7 datasets
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 196 bp overlap
ChIP MRC-5_NFD_02DPT GSE75910.NEUROG2.MRC-5_NFD_02DPT 207 bp overlap
ChIP MRC-5_NFD_05DPT GSE75910.NEUROG2.MRC-5_NFD_05DPT 279 bp overlap
ChIP MRC-5_NFD_1DPT GSE75910.NEUROG2.MRC-5_NFD_1DPT 226 bp overlap
ChIP MRC-5_N_02DPT GSE75910.NEUROG2.MRC-5_N_02DPT 254 bp overlap
ChIP MRC-5_N_05DPT GSE75910.NEUROG2.MRC-5_N_05DPT 306 bp overlap
ChIP MRC-5_N_1DPT GSE75910.NEUROG2.MRC-5_N_1DPT 498 bp overlap
NIPBL 1 dataset
ChIP GP5D GSE51234.NIPBL.GP5D 530 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 72 bp overlap
NKX3-1 1 dataset
ChIP islet ERP004003.NKX3-1.islet 184 bp overlap
NR2F2 5 datasets
ChIP K-562 ENCSR000BRS.NR2F2.K-562 215 bp overlap
ChIP liver ENCFF427MRU 421 bp overlap
ChIP liver ENCFF565JGD 491 bp overlap
ChIP liver ENCSR168SMX.NR2F2.liver 669 bp overlap
ChIP liver ENCSR338MMB.NR2F2.liver 316 bp overlap
NR2F6 1 dataset
ChIP K-562 ENCSR707QWA.NR2F6.K-562 149 bp overlap
NR3C1 2 datasets
ChIP IMR-90 ERP007093.NR3C1.IMR-90 64 bp overlap
ChIP MDA-MB-231 GSE152203.NR3C1.MDA-MB-231 53 bp overlap
OSR2 1 dataset
ChIP HEK293 GSE76494.OSR2.HEK293 204 bp overlap
PGR 2 datasets
ChIP myometrium_NP1 GSE137550.PGR.myometrium_NP1 406 bp overlap
ChIP myometrium_NP2 GSE137550.PGR.myometrium_NP2 341 bp overlap
POLR2A 3 datasets
ChIP sigmoid colon ENCFF748YVT 415 bp overlap
ChIP transverse colon ENCFF607LKE 214 bp overlap
ChIP upper lobe of left lung ENCFF199JUI 227 bp overlap
POU5F1 1 dataset
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 267 bp overlap
PRDM6 2 datasets
ChIP HEK293 ENCFF283AJL 142 bp overlap
ChIP HEK293 ENCSR892QHR.PRDM6.HEK293 222 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 417 bp overlap
RAD21 2 datasets
ChIP GP5D GSE51234.RAD21.GP5D 315 bp overlap
ChIP IMR-90 ENCSR000EFJ.RAD21.IMR-90 61 bp overlap
RARA 1 dataset
ChIP hiPSC_D3 GSE109172.RARA.hiPSC_D3 400 bp overlap
RCOR1 2 datasets
ChIP IMR-90 ENCFF644MZN 132 bp overlap
ChIP IMR-90 ENCSR000EFG.RCOR1.IMR-90 70 bp overlap
REST 6 datasets
ChIP K-562 ENCSR000BMW.REST.K-562 166 bp overlap
ChIP PFSK1 ENCSR000BOX.REST.PFSK1 89 bp overlap
ChIP colorectal-cancer_CRC121_dissociated GSE112555.REST.colorectal-cancer_CRC121_dissociated 183 bp overlap
ChIP colorectal-cancer_shCDH1 GSE112555.REST.colorectal-cancer_shCDH1 495 bp overlap
ChIP liver ENCFF240FWT 465 bp overlap
ChIP liver ENCSR867WPH.REST.liver 183 bp overlap
SIN3A 1 dataset
ChIP hiPSC_IB12 GSE106870.SIN3A.hiPSC_IB12 137 bp overlap
SKIL 1 dataset
ChIP K-562 ENCSR336DXE.SKIL.K-562 298 bp overlap
SMAD2-3 3 datasets
ChIP HGrC1_C134W-TGF GSE138496.SMAD2-3.HGrC1_C134W-TGF 123 bp overlap
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 506 bp overlap
ChIP KGN_TGF GSE138496.SMAD2-3.KGN_TGF 208 bp overlap
SMAD4 1 dataset
ChIP HGrC1_C134W-TGF GSE138496.SMAD4.HGrC1_C134W-TGF 300 bp overlap
SMARCA2 1 dataset
ChIP NPC_K755R-pos GSE122631.SMARCA2.NPC_K755R-pos 110 bp overlap
SMARCA4 5 datasets
ChIP TTC-1240 GSE124903.SMARCA4.TTC-1240 766 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCA4.TTC-1240_R377H 746 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 532 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 136 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCA4.TTC-1240_delC 766 bp overlap
SMARCB1 2 datasets
ChIP TTC-1240_R377H GSE124903.SMARCB1.TTC-1240_R377H 216 bp overlap
ChIP proliferating-human-fibroblast_ASO_Ctrl GSE131711.SMARCB1.proliferating-human-fibroblast_ASO_Ctrl 335 bp overlap
SMARCC1 4 datasets
ChIP TTC-1240 GSE124903.SMARCC1.TTC-1240 699 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCC1.TTC-1240_R377H 690 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCC1.TTC-1240_SMARCB1-FL 534 bp overlap
ChIP TTC-1240_delC GSE124903.SMARCC1.TTC-1240_delC 766 bp overlap
SMC3 3 datasets
ChIP GP5D GSE51234.SMC3.GP5D 438 bp overlap
ChIP IMR-90 ENCFF627LON 91 bp overlap
ChIP IMR-90 ENCSR000HPG.SMC3.IMR-90 64 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 527 bp overlap
SOX6 1 dataset
ChIP K-562 ENCSR788RSW.SOX6.K-562 332 bp overlap
SP7 1 dataset
ChIP HEK293 ENCSR468IJT.SP7.HEK293 356 bp overlap
SPI1 1 dataset
ChIP K-562_NABUT GSE74999.SPI1.K-562_NABUT 182 bp overlap
SS18 1 dataset
ChIP SYO-1_shSSX GSE108025.SS18.SYO-1_shSSX 100 bp overlap
SS18-SSX 1 dataset
ChIP fibroblast_23aa-SSX-Tail GSE139053.SS18-SSX.fibroblast_23aa-SSX-Tail 63 bp overlap
STAT3 2 datasets
ChIP MCF-10A ENCSR000DOQ.STAT3.MCF-10A 101 bp overlap
ChIP MCF10A-Er-Src_TAM GSE115597.STAT3.MCF10A-Er-Src_TAM 103 bp overlap
TAL1 3 datasets
ChIP CHRF28811 ERP008568.TAL1.CHRF28811 281 bp overlap
ChIP K-562 GSE107726.TAL1.K-562 572 bp overlap
ChIP K-562 ENCSR106FRG.TAL1.K-562 278 bp overlap
TBX2 1 dataset
ChIP Kelly GSE94822.TBX2.Kelly 164 bp overlap
TCF21 1 dataset
ChIP HCASMC GSE124011.TCF21.HCASMC 121 bp overlap
TCF3 2 datasets
ChIP K-562 ENCSR970OJY.TCF3.K-562 250 bp overlap
ChIP K562 ENCFF319QZT 381 bp overlap
TEAD1 1 dataset
ChIP HEK293_siE1A GSE130135.TEAD1.HEK293_siE1A 114 bp overlap
TFAP4 1 dataset
ChIP DLD-1 GSE46935.TFAP4.DLD-1 475 bp overlap
VEZF1 1 dataset
ChIP K562 ENCFF053XDV 438 bp overlap
ZBTB24 1 dataset
ChIP HCT-116 GSE111683.ZBTB24.HCT-116 154 bp overlap
ZBTB42 1 dataset
ChIP HEK293 GSE76494.ZBTB42.HEK293 187 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 459 bp overlap
ZNF711 1 dataset
ChIP HEK293T GSE145160.ZNF711.HEK293T 514 bp overlap
ZNF778 1 dataset
ChIP HEK293 GSE76494.ZNF778.HEK293 157 bp overlap