chr5 : 169,528,729 – 169,529,472
743 bp 127 TFs 2 linked genes
This 743 bp open chromatin element is linked to SPDL1 and SLIT3 and is bound by 127 transcription factors.
Linked Genes
2 genes
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
SPDL1 54.7 kb Distal Multiome
SLIT3 227.9 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr5:169,523,729 – 169,534,472
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
127 transcription factors
Source
Cell type
ATF2 4 datasets
Motif DE_36h DE_36h-ATF2_MA1632.2 10 bp overlap
Motif DE_48h DE_48h-ATF2_MA1632.2 10 bp overlap
Motif DE_60h DE_60h-ATF2_MA1632.2 10 bp overlap
Motif DE_72h DE_72h-ATF2_MA1632.2 10 bp overlap
Arid3a 1 dataset
Motif DE_60h DE_60h-Arid3a_MA0151.1 6 bp overlap
BACH2 4 datasets
Motif DE_36h DE_36h-BACH2_MA1470.2 19 bp overlap
Motif DE_48h DE_48h-BACH2_MA1470.2 19 bp overlap
Motif DE_60h DE_60h-BACH2_MA1470.2 19 bp overlap
Motif DE_72h DE_72h-BACH2_MA1470.2 19 bp overlap
BARX1 2 datasets
Motif DE_48h DE_48h-BARX1_MA0875.2 6 bp overlap
Motif DE_60h DE_60h-BARX1_MA0875.2 6 bp overlap
BRD4 1 dataset
ChIP HAP1 GSE108387.BRD4.HAP1 517 bp overlap
BSX 2 datasets
Motif DE_48h DE_48h-BSX_MA0876.2 6 bp overlap
Motif DE_60h DE_60h-BSX_MA0876.2 6 bp overlap
CEBPB 1 dataset
ChIP HFOB_DIFF GSE82295.CEBPB.HFOB_DIFF 272 bp overlap
Crx 1 dataset
Motif DE_60h DE_60h-Crx_MA0467.3 6 bp overlap
DLX1 2 datasets
Motif DE_48h DE_48h-DLX1_MA0879.3 6 bp overlap
Motif DE_60h DE_60h-DLX1_MA0879.3 6 bp overlap
DLX6 2 datasets
Motif DE_48h DE_48h-DLX6_MA0882.2 6 bp overlap
Motif DE_60h DE_60h-DLX6_MA0882.2 6 bp overlap
DUX4 5 datasets
Motif DE_36h DE_36h-DUX4_MA0468.1 11 bp overlap
Motif DE_48h DE_48h-DUX4_MA0468.1 11 bp overlap
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
Motif DE_72h DE_72h-DUX4_MA0468.1 11 bp overlap
ChIP WA01 GSE94322.DUX4.WA01 194 bp overlap
Dlx3 2 datasets
Motif DE_48h DE_48h-Dlx3_MA0880.2 6 bp overlap
Motif DE_60h DE_60h-Dlx3_MA0880.2 6 bp overlap
Dlx4 2 datasets
Motif DE_48h DE_48h-Dlx4_MA0881.2 6 bp overlap
Motif DE_60h DE_60h-Dlx4_MA0881.2 6 bp overlap
Dmbx1 1 dataset
Motif DE_60h DE_60h-Dmbx1_MA0883.2 10 bp overlap
EN2 2 datasets
Motif DE_48h DE_48h-EN2_MA0642.3 7 bp overlap
Motif DE_60h DE_60h-EN2_MA0642.3 7 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 199 bp overlap
ERF::SREBF2 5 datasets
Motif DE_36h DE_36h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_48h DE_48h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_48h DE_48h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_60h DE_60h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_72h DE_72h-ERFSREBF2_MA1939.2 16 bp overlap
ERG 1 dataset
ChIP aortic-endothelial-cell_D40 GSE139377.ERG.aortic-endothelial-cell_D40 204 bp overlap
ETV2 1 dataset
Motif DE_48h DE_48h-ETV2_MA0762.2 9 bp overlap
FIGLA 4 datasets
Motif DE_36h DE_36h-FIGLA_MA0820.2 6 bp overlap
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
Motif DE_72h DE_72h-FIGLA_MA0820.2 6 bp overlap
FOS 4 datasets
Motif DE_36h DE_36h-FOS_MA1951.2 13 bp overlap
Motif DE_48h DE_48h-FOS_MA1951.2 13 bp overlap
Motif DE_60h DE_60h-FOS_MA1951.2 13 bp overlap
Motif DE_72h DE_72h-FOS_MA1951.2 13 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 355 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 743 bp overlap
ChIP DE DE-FOXA2-2 642 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 341 bp overlap
FOXC1 1 dataset
Motif DE_48h DE_48h-FOXC1_MA0032.2 11 bp overlap
FOXC2 1 dataset
Motif DE_48h DE_48h-FOXC2_MA0846.2 11 bp overlap
FOXD2 1 dataset
Motif DE_48h DE_48h-FOXD2_MA0847.4 11 bp overlap
FOXE1 1 dataset
Motif DE_48h DE_48h-FOXE1_MA1487.3 12 bp overlap
FOXS1 1 dataset
Motif DE_48h DE_48h-FOXS1_MA2118.1 8 bp overlap
Foxl2 1 dataset
Motif DE_48h DE_48h-Foxl2_MA1607.2 10 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 612 bp overlap
ChIP DE DE-GATA4-2 712 bp overlap
ChIP foregut GSE117136.GATA4.foregut 555 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 594 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 482 bp overlap
GATA6 14 datasets
ChIP AGS GSE51705.GATA6.AGS 156 bp overlap
ChIP DE DE-GATA6-1 650 bp overlap
ChIP DE DE-GATA6-2 711 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 585 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 661 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 643 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 705 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 741 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 692 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 358 bp overlap
ChIP foregut GSE117136.GATA6.foregut 673 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 602 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 480 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 479 bp overlap
GBX1 2 datasets
Motif DE_48h DE_48h-GBX1_MA0889.2 7 bp overlap
Motif DE_60h DE_60h-GBX1_MA0889.2 7 bp overlap
GBX2 2 datasets
Motif DE_48h DE_48h-GBX2_MA0890.2 6 bp overlap
Motif DE_60h DE_60h-GBX2_MA0890.2 6 bp overlap
GRHL1 2 datasets
Motif DE_48h DE_48h-GRHL1_MA0647.2 10 bp overlap
Motif DE_60h DE_60h-GRHL1_MA0647.2 10 bp overlap
GSC 1 dataset
Motif DE_60h DE_60h-GSC_MA0648.2 6 bp overlap
GSC2 1 dataset
Motif DE_60h DE_60h-GSC2_MA0891.2 6 bp overlap
HESX1 2 datasets
Motif DE_48h DE_48h-HESX1_MA0894.2 6 bp overlap
Motif DE_60h DE_60h-HESX1_MA0894.2 6 bp overlap
HOXA7 2 datasets
Motif DE_48h DE_48h-HOXA7_MA1498.3 6 bp overlap
Motif DE_60h DE_60h-HOXA7_MA1498.3 6 bp overlap
Hand1 1 dataset
Motif DE_48h DE_48h-Hand1_MA2123.1 9 bp overlap
Hnf1A 1 dataset
Motif DE_48h DE_48h-Hnf1A_MA1991.2 10 bp overlap
IKZF1 1 dataset
Motif DE_48h DE_48h-IKZF1_MA1508.2 8 bp overlap
IKZF2 1 dataset
Motif DE_48h DE_48h-IKZF2_MA2326.1 6 bp overlap
IRF7 1 dataset
Motif DE_48h DE_48h-IRF7_MA0772.2 13 bp overlap
ISL2 1 dataset
Motif DE_48h DE_48h-ISL2_MA0914.2 6 bp overlap
Irf1 1 dataset
Motif DE_48h DE_48h-Irf1_MA0050.4 11 bp overlap
JUN 3 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 561 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 420 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 602 bp overlap
KDM1A 3 datasets
ChIP pancreatic-progenitor_GT GSE104840.KDM1A.pancreatic-progenitor_GT 350 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 213 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 413 bp overlap
LBX1 2 datasets
Motif DE_48h DE_48h-LBX1_MA0618.2 7 bp overlap
Motif DE_60h DE_60h-LBX1_MA0618.2 7 bp overlap
LBX2 2 datasets
Motif DE_48h DE_48h-LBX2_MA0699.2 6 bp overlap
Motif DE_60h DE_60h-LBX2_MA0699.2 6 bp overlap
LHX2 2 datasets
Motif DE_48h DE_48h-LHX2_MA0700.3 6 bp overlap
Motif DE_60h DE_60h-LHX2_MA0700.3 6 bp overlap
LHX6 1 dataset
Motif DE_48h DE_48h-LHX6_MA0658.2 8 bp overlap
LHX9 2 datasets
Motif DE_48h DE_48h-LHX9_MA0701.3 7 bp overlap
Motif DE_60h DE_60h-LHX9_MA0701.3 7 bp overlap
MGA::EVX1 2 datasets
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
MSX1 2 datasets
Motif DE_48h DE_48h-MSX1_MA0666.3 6 bp overlap
Motif DE_60h DE_60h-MSX1_MA0666.3 6 bp overlap
MSX2 2 datasets
Motif DE_48h DE_48h-MSX2_MA0708.3 6 bp overlap
Motif DE_60h DE_60h-MSX2_MA0708.3 6 bp overlap
Msx3 2 datasets
Motif DE_48h DE_48h-Msx3_MA0709.2 6 bp overlap
Motif DE_60h DE_60h-Msx3_MA0709.2 6 bp overlap
NANOG 2 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 393 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 301 bp overlap
NBN 1 dataset
ChIP MCF-10A GSE93038.NBN.MCF-10A 285 bp overlap
NFIA 2 datasets
Motif DE_48h DE_48h-NFIA_MA0670.2 6 bp overlap
Motif DE_60h DE_60h-NFIA_MA0670.2 6 bp overlap
NFIX 2 datasets
Motif DE_48h DE_48h-NFIX_MA0671.2 6 bp overlap
Motif DE_60h DE_60h-NFIX_MA0671.2 6 bp overlap
NKX2-2 2 datasets
Motif DE_48h DE_48h-NKX2-2_MA1645.2 8 bp overlap
Motif DE_60h DE_60h-NKX2-2_MA1645.2 8 bp overlap
NKX2-3 1 dataset
Motif DE_48h DE_48h-NKX2-3_MA0672.2 8 bp overlap
NKX2-4 1 dataset
Motif DE_48h DE_48h-NKX2-4_MA2003.2 8 bp overlap
NKX2-8 1 dataset
Motif DE_48h DE_48h-NKX2-8_MA0673.2 8 bp overlap
NR1D1 4 datasets
Motif DE_36h DE_36h-NR1D1_MA1531.2 14 bp overlap
Motif DE_48h DE_48h-NR1D1_MA1531.2 14 bp overlap
Motif DE_60h DE_60h-NR1D1_MA1531.2 14 bp overlap
Motif DE_72h DE_72h-NR1D1_MA1531.2 14 bp overlap
NR1D2 4 datasets
Motif DE_36h DE_36h-NR1D2_MA1532.2 15 bp overlap
Motif DE_48h DE_48h-NR1D2_MA1532.2 15 bp overlap
Motif DE_60h DE_60h-NR1D2_MA1532.2 15 bp overlap
Motif DE_72h DE_72h-NR1D2_MA1532.2 15 bp overlap
Nkx3-1 1 dataset
Motif DE_48h DE_48h-Nkx3-1_MA0124.3 7 bp overlap
Nkx3-2 1 dataset
Motif DE_48h DE_48h-Nkx3-2_MA0122.4 10 bp overlap
Nobox 2 datasets
Motif DE_48h DE_48h-Nobox_MA0125.2 6 bp overlap
Motif DE_60h DE_60h-Nobox_MA0125.2 6 bp overlap
Nr2F6 4 datasets
Motif DE_36h DE_36h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_48h DE_48h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_60h DE_60h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_72h DE_72h-Nr2F6_MA0728.1 15 bp overlap
ONECUT1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 409 bp overlap
OTX1 1 dataset
Motif DE_60h DE_60h-OTX1_MA0711.2 6 bp overlap
PDX1 3 datasets
ChIP hESC GSE58685.PDX1.hESC 156 bp overlap
ChIP hiPSC GSE125768.PDX1.hiPSC 456 bp overlap
ChIP pancreatic-progenitor GSE117136.PDX1.pancreatic-progenitor 288 bp overlap
PITX1 1 dataset
Motif DE_60h DE_60h-PITX1_MA0682.3 6 bp overlap
PITX2 1 dataset
Motif DE_60h DE_60h-PITX2_MA1547.2 8 bp overlap
PITX3 1 dataset
Motif DE_60h DE_60h-PITX3_MA0714.2 6 bp overlap
POU2F3 1 dataset
Motif DE_48h DE_48h-POU2F3_MA0627.3 9 bp overlap
POU5F1 1 dataset
Motif DE_48h DE_48h-POU5F1_MA1115.2 7 bp overlap
PRRX2 2 datasets
Motif DE_48h DE_48h-PRRX2_MA0075.4 7 bp overlap
Motif DE_60h DE_60h-PRRX2_MA0075.4 7 bp overlap
RAX 2 datasets
Motif DE_48h DE_48h-RAX_MA0718.2 6 bp overlap
Motif DE_60h DE_60h-RAX_MA0718.2 6 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 146 bp overlap
RHOXF1 1 dataset
Motif DE_60h DE_60h-RHOXF1_MA0719.2 6 bp overlap
RNF2 1 dataset
ChIP fibroblast GSE139053.RNF2.fibroblast 151 bp overlap
RUNX3 4 datasets
Motif DE_36h DE_36h-RUNX3_MA0684.3 8 bp overlap
Motif DE_48h DE_48h-RUNX3_MA0684.3 8 bp overlap
Motif DE_60h DE_60h-RUNX3_MA0684.3 8 bp overlap
Motif DE_72h DE_72h-RUNX3_MA0684.3 8 bp overlap
Rarb 5 datasets
Motif DE_36h DE_36h-Rarb_MA0857.1 16 bp overlap
Motif DE_48h DE_48h-Rarb_MA0857.1 16 bp overlap
Motif DE_48h DE_48h-Rarb_MA0858.1 17 bp overlap
Motif DE_60h DE_60h-Rarb_MA0857.1 16 bp overlap
Motif DE_72h DE_72h-Rarb_MA0857.1 16 bp overlap
SMAD2 1 dataset
Motif DE_48h DE_48h-SMAD2_MA1964.2 6 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 680 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 714 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 611 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 488 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 639 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 567 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 517 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 603 bp overlap
SMAD4 2 datasets
ChIP Caco-2 GSE112946.SMAD4.Caco-2 169 bp overlap
ChIP endoderm GSE29422.SMAD4.endoderm 215 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 409 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 301 bp overlap
SOX14 1 dataset
Motif DE_48h DE_48h-SOX14_MA1562.2 9 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 305 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 604 bp overlap
SOX8 1 dataset
Motif DE_48h DE_48h-SOX8_MA0868.3 7 bp overlap
SP5 1 dataset
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
SREBF1 4 datasets
Motif DE_36h DE_36h-SREBF1_MA0829.3 10 bp overlap
Motif DE_48h DE_48h-SREBF1_MA0829.3 10 bp overlap
Motif DE_60h DE_60h-SREBF1_MA0829.3 10 bp overlap
Motif DE_72h DE_72h-SREBF1_MA0829.3 10 bp overlap
SRY 1 dataset
Motif DE_48h DE_48h-SRY_MA0084.2 7 bp overlap
Sox5 1 dataset
Motif DE_48h DE_48h-Sox5_MA0087.3 8 bp overlap
Spz1 4 datasets
Motif DE_36h DE_36h-Spz1_MA0111.1 11 bp overlap
Motif DE_48h DE_48h-Spz1_MA0111.1 11 bp overlap
Motif DE_60h DE_60h-Spz1_MA0111.1 11 bp overlap
Motif DE_72h DE_72h-Spz1_MA0111.1 11 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 190 bp overlap
Tfcp2l1 1 dataset
Motif DE_60h DE_60h-Tfcp2l1_MA0145.2 14 bp overlap
Vdr 4 datasets
Motif DE_36h DE_36h-Vdr_MA0693.4 7 bp overlap
Motif DE_48h DE_48h-Vdr_MA0693.4 7 bp overlap
Motif DE_60h DE_60h-Vdr_MA0693.4 7 bp overlap
Motif DE_72h DE_72h-Vdr_MA0693.4 7 bp overlap
ZBTB24 1 dataset
Motif DE_48h DE_48h-ZBTB24_MA2330.1 10 bp overlap
ZEB1 4 datasets
Motif DE_36h DE_36h-ZEB1_MA0103.4 6 bp overlap
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
Motif DE_72h DE_72h-ZEB1_MA0103.4 6 bp overlap
ZFP14 1 dataset
Motif DE_48h DE_48h-ZFP14_MA1972.1 15 bp overlap
ZFP42 1 dataset
Motif DE_36h DE_36h-ZFP42_MA1651.2 13 bp overlap
ZNF135 1 dataset
Motif DE_48h DE_48h-ZNF135_MA1587.1 14 bp overlap
ZNF16 2 datasets
Motif DE_48h DE_48h-ZNF16_MA1654.2 21 bp overlap
Motif DE_72h DE_72h-ZNF16_MA1654.2 21 bp overlap
ZNF189 1 dataset
Motif DE_48h DE_48h-ZNF189_MA1725.2 9 bp overlap
ZNF213 1 dataset
Motif DE_36h DE_36h-ZNF213_MA2121.1 12 bp overlap
ZNF263 1 dataset
Motif DE_48h DE_48h-ZNF263_MA0528.3 7 bp overlap
ZNF28 1 dataset
ChIP HEK293T GSE78099.ZNF28.HEK293T 189 bp overlap
ZNF282 1 dataset
Motif DE_48h DE_48h-ZNF282_MA1154.2 15 bp overlap
ZNF341 1 dataset
Motif DE_48h DE_48h-ZNF341_MA1655.2 8 bp overlap
ZNF449 1 dataset
Motif DE_36h DE_36h-ZNF449_MA1656.2 10 bp overlap
Zic3 1 dataset
Motif DE_48h DE_48h-Zic3_MA0697.3 7 bp overlap