chr6 : 28,357,204 – 28,357,431
227 bp 129 TFs 2 linked genes
This 227 bp open chromatin element is linked to ZSCAN31 and ZKSCAN3 and is bound by 129 transcription factors.
Linked Genes
2 genes
Distance
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
ZSCAN31 933 bp At TSS Proximity
ZKSCAN3 7.3 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr6:28,352,204 – 28,362,431
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
129 transcription factors
Source
Cell type
ARID1B 2 datasets
ChIP K-562 ENCSR822CCM.ARID1B.K-562 206 bp overlap
ChIP K562 ENCFF938UXQ 227 bp overlap
ARNT 1 dataset
ChIP K-562 ENCSR155KHM.ARNT.K-562 227 bp overlap
ARNTL 3 datasets
ChIP U2OS GSE130602.ARNTL.U2OS 147 bp overlap
ChIP U2OS_DMSO GSE130506.ARNTL.U2OS_DMSO 147 bp overlap
ChIP U2OS_cordycepin GSE130506.ARNTL.U2OS_cordycepin 123 bp overlap
ATF1 1 dataset
ChIP K562 ENCFF469GPI 227 bp overlap
ATF3 2 datasets
ChIP K-562 ENCSR028UIU.ATF3.K-562 190 bp overlap
ChIP K562 ENCFF604FPV 227 bp overlap
ATF7 1 dataset
ChIP K562 ENCFF308SKS 227 bp overlap
BACH1 6 datasets
ChIP AsPC-1 GSE124406.BACH1.AsPC-1 183 bp overlap
ChIP K-562 ENCSR740NPG.BACH1.K-562 227 bp overlap
ChIP K562 ENCFF419VIM 227 bp overlap
ChIP K562 ENCFF419VIM 227 bp overlap
ChIP K562 ENCFF990JHI 100 bp overlap
ChIP SW1990 GSE124406.BACH1.SW1990 185 bp overlap
BAF155 1 dataset
ChIP VCaP_shARID1A GSE110655.BAF155.VCaP_shARID1A 193 bp overlap
BCOR 2 datasets
ChIP K-562 ENCSR808AKZ.BCOR.K-562 69 bp overlap
ChIP K562 ENCFF343XWA 190 bp overlap
BHLHE40 2 datasets
ChIP IMR-90 ENCFF312JYK 107 bp overlap
ChIP IMR-90 ENCSR957KYB.BHLHE40.IMR-90 59 bp overlap
BRD2 6 datasets
ChIP MDA-MB-231_BAZ2AsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2AsiRNA 227 bp overlap
ChIP MDA-MB-231_BAZ2BsiRNA GSE116879.BRD2.MDA-MB-231_BAZ2BsiRNA 227 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD2.MDA-MB-231_MGSK2801 227 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD2.MDA-MB-231_MGSK2801_rDNA 227 bp overlap
ChIP MDA-MB-231_NTsiRNA GSE116879.BRD2.MDA-MB-231_NTsiRNA 227 bp overlap
ChIP SK-MEL-147 GSE94488.BRD2.SK-MEL-147 227 bp overlap
BRD4 10 datasets
ChIP IMR-90_QUIES GSE74238.BRD4.IMR-90_QUIES 119 bp overlap
ChIP MDA-MB-231_DMSO GSE116879.BRD4.MDA-MB-231_DMSO 86 bp overlap
ChIP MDA-MB-231_DMSO_rDNA GSE116879.BRD4.MDA-MB-231_DMSO_rDNA 86 bp overlap
ChIP MDA-MB-231_JQ1-neg_L GSE136151.BRD4.MDA-MB-231_JQ1-neg_L 196 bp overlap
ChIP MDA-MB-231_JQ1-neg_S_3f GSE136151.BRD4.MDA-MB-231_JQ1-neg_S_3f 227 bp overlap
ChIP MDA-MB-231_MGSK2801 GSE116879.BRD4.MDA-MB-231_MGSK2801 206 bp overlap
ChIP MDA-MB-231_MGSK2801_rDNA GSE116879.BRD4.MDA-MB-231_MGSK2801_rDNA 206 bp overlap
ChIP SUM159PT_30nMbortezomib_8h GSE87418.BRD4.SUM159PT_30nMbortezomib_8h 142 bp overlap
ChIP SUM159PT_shMYC_DMSO_48h GSE87418.BRD4.SUM159PT_shMYC_DMSO_48h 226 bp overlap
ChIP SUM159PT_shMYC_dox_48h GSE87418.BRD4.SUM159PT_shMYC_dox_48h 196 bp overlap
BRD9 2 datasets
ChIP K562 ENCFF480JXZ 142 bp overlap
ChIP K562 ENCFF480JXZ 227 bp overlap
CAVIN1 1 dataset
ChIP K562 ENCFF731YBD 223 bp overlap
CBX3 1 dataset
ChIP K562 ENCFF410AQU 91 bp overlap
CBX5 2 datasets
ChIP K-562 ENCSR272JAT.CBX5.K-562 55 bp overlap
ChIP K562 ENCFF188CYP 115 bp overlap
CEBPZ 2 datasets
ChIP K-562 ENCSR618GDK.CEBPZ.K-562 77 bp overlap
ChIP K562 ENCFF909BYC 136 bp overlap
CREB3 3 datasets
ChIP K-562 ENCSR093FKD.CREB3.K-562 227 bp overlap
ChIP K562 ENCFF985QJI 227 bp overlap
ChIP K562 ENCFF985QJI 196 bp overlap
CREBBP 1 dataset
ChIP fibroblast_senescent GSE106146.CREBBP.fibroblast_senescent 101 bp overlap
CREM 1 dataset
ChIP K562 ENCFF180STA 96 bp overlap
CTCF 1 dataset
ChIP HCT-116 GSE131606.CTCF.HCT-116 178 bp overlap
CUX1 1 dataset
ChIP K562 ENCFF902MYN 227 bp overlap
CXXC5 1 dataset
ChIP K562 ENCFF497CZN 227 bp overlap
DMBX1 1 dataset
ChIP K562 ENCFF972HXB 200 bp overlap
DPF2 1 dataset
ChIP K562 ENCFF775HUO 69 bp overlap
E2F1 1 dataset
ChIP MDA-MB-231 GSE95303.E2F1.MDA-MB-231 227 bp overlap
E4F1 1 dataset
ChIP K-562 ENCSR731LHZ.E4F1.K-562 212 bp overlap
EHMT2 1 dataset
ChIP K-562 ENCSR175EOM.EHMT2.K-562 186 bp overlap
ELF1 1 dataset
ChIP K-562 ENCSR000BMD.ELF1.K-562 57 bp overlap
EMSY 2 datasets
ChIP K562 ENCFF511ZZZ 164 bp overlap
ChIP K562 ENCFF511ZZZ 227 bp overlap
EP300 3 datasets
ChIP fibroblast_senescent GSE106146.EP300.fibroblast_senescent 218 bp overlap
ChIP pulmonary-artery_endothelial-cell_siCtrl GSE89786.EP300.pulmonary-artery_endothelial-cell_siCtrl 173 bp overlap
ChIP pulmonary-artery_endothelial-cell_siPFKFB3 GSE89786.EP300.pulmonary-artery_endothelial-cell_siPFKFB3 144 bp overlap
ERG 2 datasets
ChIP HUVEC-C GSE124891.ERG.HUVEC-C 127 bp overlap
ChIP VCaP_shARID1A GSE110655.ERG.VCaP_shARID1A 203 bp overlap
ETS1 7 datasets
ChIP HUVEC-C GSE41166.ETS1.HUVEC-C 120 bp overlap
ChIP HUVEC-C_12h GSE93030.ETS1.HUVEC-C_12h 151 bp overlap
ChIP HUVEC-C_4h GSE93030.ETS1.HUVEC-C_4h 82 bp overlap
ChIP HUVEC-C_VEGF_12H GSE41166.ETS1.HUVEC-C_VEGF_12H 185 bp overlap
ChIP HUVEC-C_VEGF_12h GSE109625.ETS1.HUVEC-C_VEGF_12h 151 bp overlap
ChIP HUVEC-C_VEGF_4H GSE41166.ETS1.HUVEC-C_VEGF_4H 102 bp overlap
ChIP HUVEC-C_VEGF_4h GSE109625.ETS1.HUVEC-C_VEGF_4h 82 bp overlap
ETV1 2 datasets
ChIP K-562 ENCSR277DMR.ETV1.K-562 138 bp overlap
ChIP K562 ENCFF389WTI 176 bp overlap
FLI1 1 dataset
ChIP HUVEC-C GSE109695.FLI1.HUVEC-C 147 bp overlap
FOS 2 datasets
ChIP IMR-90 ENCFF179EDA 137 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 130 bp overlap
FOSL1 1 dataset
ChIP MDA-MB-231 GSE95303.FOSL1.MDA-MB-231 137 bp overlap
FOXA2 2 datasets
ChIP CFPAC-1 GSE119930.FOXA2.CFPAC-1 160 bp overlap
ChIP DE DE-FOXA2-1 210 bp overlap
FOXA3 2 datasets
ChIP K562 ENCFF348SOM 227 bp overlap
ChIP K562 ENCFF781VSC 182 bp overlap
FOXK2 4 datasets
ChIP K-562 ENCSR508DQA.FOXK2.K-562 109 bp overlap
ChIP K-562 ENCSR302AWT.FOXK2.K-562 227 bp overlap
ChIP K562 ENCFF245WKP 219 bp overlap
ChIP K562 ENCFF851PFH 219 bp overlap
GABPA 1 dataset
ChIP K562 ENCFF139LXS 227 bp overlap
GABPB1 2 datasets
ChIP K-562 ENCSR138YYY.GABPB1.K-562 227 bp overlap
ChIP K562 ENCFF015GDS 227 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 227 bp overlap
ChIP DE DE-GATA4-2 227 bp overlap
GATA6 4 datasets
ChIP DE DE-GATA6-1 227 bp overlap
ChIP DE DE-GATA6-2 227 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 227 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 227 bp overlap
HDAC2 2 datasets
ChIP K-562 ENCSR893WSB.HDAC2.K-562 227 bp overlap
ChIP K-562 ENCSR075HTM.HDAC2.K-562 226 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR197ALX.HDGF.K-562 227 bp overlap
HIF1A 2 datasets
ChIP HUVEC-C_HYPOX GSE39089.HIF1A.HUVEC-C_HYPOX 227 bp overlap
ChIP U2OS_DMOG GSE85096.HIF1A.U2OS_DMOG 84 bp overlap
HINFP 1 dataset
ChIP K562 ENCFF361QXJ 202 bp overlap
HLF 1 dataset
ChIP HepG2 ENCFF854JLR 60 bp overlap
IRF1 1 dataset
ChIP K-562 ENCSR854MCV.IRF1.K-562 227 bp overlap
JUN 3 datasets
ChIP K562 ENCFF372VWH 151 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 94 bp overlap
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 79 bp overlap
JUND 3 datasets
ChIP K-562 ENCSR000EGN.JUND.K-562 61 bp overlap
ChIP K562 ENCFF336RCR 209 bp overlap
ChIP K562 ENCFF830LVJ 112 bp overlap
KDM1A 1 dataset
ChIP K562 ENCFF128TYE 227 bp overlap
KLF13 1 dataset
ChIP K562 ENCFF738YZC 209 bp overlap
KLF6 2 datasets
ChIP K-562 ENCSR297CGF.KLF6.K-562 195 bp overlap
ChIP K562 ENCFF948QSP 227 bp overlap
L3MBTL2 2 datasets
ChIP K-562 ENCSR530XQI.L3MBTL2.K-562 227 bp overlap
ChIP K562 ENCFF320EQC 227 bp overlap
LEF1 1 dataset
ChIP K562 ENCFF198WCP 227 bp overlap
MAFB 1 dataset
ChIP islet ERP004003.MAFB.islet 64 bp overlap
MAFF 2 datasets
ChIP K-562 ENCSR000EGI.MAFF.K-562 98 bp overlap
ChIP K562 ENCFF071YKK 125 bp overlap
MAFG 2 datasets
ChIP K-562 ENCSR818DQV.MAFG.K-562 227 bp overlap
ChIP K562 ENCFF455EEO 206 bp overlap
MAFK 4 datasets
ChIP H1 ENCFF854XWE 102 bp overlap
ChIP IMR-90 ENCFF336DHZ 138 bp overlap
ChIP K-562 ENCSR000EGX.MAFK.K-562 88 bp overlap
ChIP K562 ENCFF380WHM 137 bp overlap
MAX 1 dataset
ChIP K562 ENCFF110LJS 141 bp overlap
MED1 1 dataset
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 184 bp overlap
MEF2D 1 dataset
ChIP K562 ENCFF392LDT 227 bp overlap
MGA 1 dataset
ChIP K562 ENCFF140CEX 227 bp overlap
MNT 1 dataset
ChIP K562 ENCFF820IGH 227 bp overlap
MTA3 1 dataset
ChIP K-562 ENCSR914NEI.MTA3.K-562 227 bp overlap
MYC 1 dataset
ChIP endothelial_umbilical-vein ENCSR000DLU.MYC.endothelial_umbilical-vein 53 bp overlap
NCOR1 1 dataset
ChIP K-562 ENCSR798ILC.NCOR1.K-562 200 bp overlap
NFE2 5 datasets
ChIP K-562 ENCSR552YGL.NFE2.K-562 185 bp overlap
ChIP K-562 ENCSR000FCC.NFE2.K-562 145 bp overlap
ChIP K562 ENCFF047YKA 180 bp overlap
ChIP K562 ENCFF163BSI 76 bp overlap
ChIP erythroid GSE125753.NFE2.erythroid 64 bp overlap
NFE2L1 2 datasets
ChIP K-562 ENCSR632SHZ.NFE2L1.K-562 185 bp overlap
ChIP K562 ENCFF138UHK 130 bp overlap
NFE2L2 7 datasets
ChIP A-549 ENCSR584GHV.NFE2L2.A-549 85 bp overlap
ChIP A549 ENCFF474YMB 118 bp overlap
ChIP Hep-G2 ENCSR488EES.NFE2L2.Hep-G2 68 bp overlap
ChIP HepG2 ENCFF178DRC 113 bp overlap
ChIP IMR-90 ENCFF059WEE 86 bp overlap
ChIP IMR-90 ENCSR197WGI.NFE2L2.IMR-90 204 bp overlap
ChIP K-562 ENCSR485GKE.NFE2L2.K-562 200 bp overlap
NFRKB 1 dataset
ChIP K-562 ENCSR657EOF.NFRKB.K-562 227 bp overlap
NR1H2 1 dataset
ChIP K562 ENCFF386VZB 172 bp overlap
NR2C1 2 datasets
ChIP K-562 ENCSR178DEG.NR2C1.K-562 168 bp overlap
ChIP K562 ENCFF568JLK 219 bp overlap
NR4A1 2 datasets
ChIP K-562 ENCSR130PDE.NR4A1.K-562 227 bp overlap
ChIP K562 ENCFF998LHF 227 bp overlap
POLR2A 3 datasets
ChIP Panc1 ENCFF290KAB 227 bp overlap
ChIP endothelial cell of umbilical vein ENCFF467WJF 227 bp overlap
ChIP stomach ENCFF820WZN 174 bp overlap
PTTG1 1 dataset
ChIP K562 ENCFF017KRM 158 bp overlap
PYGO2 1 dataset
ChIP K562 ENCFF458IIA 149 bp overlap
RELA 1 dataset
ChIP aortic-endothelial-cell_IL1B_D8 GSE139377.RELA.aortic-endothelial-cell_IL1B_D8 70 bp overlap
RFX5 2 datasets
ChIP IMR-90 ENCFF886KPO 96 bp overlap
ChIP IMR-90 ENCSR000EFD.RFX5.IMR-90 58 bp overlap
RNF2 1 dataset
ChIP K562 ENCFF653BQJ 227 bp overlap
RUVBL2 1 dataset
ChIP U2OS GSE130602.RUVBL2.U2OS 227 bp overlap
SETDB1 2 datasets
ChIP K-562 ENCSR000EWI.SETDB1.K-562 117 bp overlap
ChIP K562 ENCFF745PAW 174 bp overlap
SMAD2-3 1 dataset
ChIP HUVEC-C_TGFB1 GSE134556.SMAD2-3.HUVEC-C_TGFB1 207 bp overlap
SMAD3 1 dataset
ChIP K-562 ENCSR376XAV.SMAD3.K-562 125 bp overlap
SMAD4 1 dataset
ChIP K562 ENCFF628RBP 227 bp overlap
SMARCE1 1 dataset
ChIP K562 ENCFF690CFF 227 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 227 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 227 bp overlap
SREBF1 1 dataset
ChIP K562 ENCFF441TTT 227 bp overlap
SRF 2 datasets
ChIP K562 ENCFF766EOO 170 bp overlap
ChIP K562 ENCFF766EOO 227 bp overlap
STAT3 1 dataset
ChIP MDA-MB-231_EtOH GSE85579.STAT3.MDA-MB-231_EtOH 69 bp overlap
TEAD1 1 dataset
ChIP K-562 ENCSR591ASD.TEAD1.K-562 134 bp overlap
TEAD4 1 dataset
ChIP K562 ENCFF843TII 227 bp overlap
TFDP1 1 dataset
ChIP K562 ENCFF794ZXJ 227 bp overlap
TP53 2 datasets
ChIP IMR-90_NUT3A GSE58740.TP53.IMR-90_NUT3A 169 bp overlap
ChIP MDA-MB-231 GSE95303.TP53.MDA-MB-231 224 bp overlap
TRIM24 2 datasets
ChIP K-562 ENCSR907MZR.TRIM24.K-562 167 bp overlap
ChIP K562 ENCFF616RIL 198 bp overlap
TRIM28 6 datasets
ChIP K-562 ENCSR000EVY.TRIM28.K-562 227 bp overlap
ChIP K-562 ENCSR474CVP.TRIM28.K-562 198 bp overlap
ChIP K562 ENCFF172UPN 159 bp overlap
ChIP K562 ENCFF671NCI 227 bp overlap
ChIP K562 ENCFF671NCI 227 bp overlap
ChIP hESC GSE115387.TRIM28.hESC 147 bp overlap
YY1 2 datasets
ChIP K-562 ENCSR000BMH.YY1.K-562 104 bp overlap
ChIP K562 ENCFF660QRE 149 bp overlap
ZBTB12 1 dataset
ChIP K562 ENCFF933CVM 227 bp overlap
ZBTB40 1 dataset
ChIP K562 ENCFF337GJB 227 bp overlap
ZBTB9 1 dataset
ChIP K562 ENCFF233EFX 207 bp overlap
ZEB2 2 datasets
ChIP K562 ENCFF795CMH 227 bp overlap
ChIP K562 ENCFF975RXS 196 bp overlap
ZFP36 1 dataset
ChIP K562 ENCFF255RZG 105 bp overlap
ZFX 3 datasets
ChIP K562 ENCFF169LZT 227 bp overlap
ChIP K562 ENCFF169LZT 227 bp overlap
ChIP K562 ENCFF169LZT 158 bp overlap
ZMYND8 1 dataset
ChIP MDA-MB-231 GSE108833.ZMYND8.MDA-MB-231 68 bp overlap
ZNF134 1 dataset
ChIP K562 ENCFF502NWS 227 bp overlap
ZNF143 1 dataset
ChIP K562 ENCFF554TVF 227 bp overlap
ZNF215 1 dataset
ChIP K562 ENCFF317MOH 183 bp overlap
ZNF24 2 datasets
ChIP K-562 ENCSR695EQB.ZNF24.K-562 227 bp overlap
ChIP K562 ENCFF877JCX 227 bp overlap
ZNF280A 1 dataset
ChIP K562 ENCFF706EWX 104 bp overlap
ZNF281 1 dataset
ChIP K562 ENCFF594VNM 227 bp overlap
ZNF282 2 datasets
ChIP K562 ENCFF657WOV 94 bp overlap
ChIP K562 ENCFF657WOV 227 bp overlap
ZNF311 1 dataset
ChIP K562 ENCFF986QSP 183 bp overlap
ZNF316 1 dataset
ChIP K562 ENCFF838QCD 177 bp overlap
ZNF317 1 dataset
ChIP K562 ENCFF896LCF 171 bp overlap
ZNF395 1 dataset
ChIP K562 ENCFF464EIT 227 bp overlap
ZNF449 1 dataset
ChIP K562 ENCFF038VEZ 227 bp overlap
ZNF592 2 datasets
ChIP K-562 ENCSR249BHQ.ZNF592.K-562 227 bp overlap
ChIP K562 ENCFF547OSS 227 bp overlap
ZNF639 1 dataset
ChIP K562 ENCFF271FQR 227 bp overlap
ZNF740 1 dataset
ChIP K562 ENCFF447IXE 227 bp overlap
ZNF83 2 datasets
ChIP K-562 ENCSR257XVY.ZNF83.K-562 164 bp overlap
ChIP K562 ENCFF340RTV 227 bp overlap