chr6 : 84,086,644 – 84,088,326
1,682 bp 191 TFs 3 linked genes
This 1.7 kb open chromatin element is linked to MRAP2, CEP162, and CYB5R4 and is bound by 191 transcription factors.
Linked Genes
3 genes
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
MRAP2 53.5 kb Distal Multiome
CEP162 140.3 kb Distal Multiome
CYB5R4 227.6 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr6:84,081,644 – 84,093,326
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
191 transcription factors
Source
Cell type
AR 2 datasets
ChIP LNCaP-abl GSE39459.AR.LNCaP-abl 106 bp overlap
ChIP VCaP_DHT24H_SHFOXP1 GSE58428.AR.VCaP_DHT24H_SHFOXP1 184 bp overlap
ARID1A 3 datasets
ChIP RMG-I GSE104545.ARID1A.RMG-I 427 bp overlap
ChIP RMG-I GSE120058.ARID1A.RMG-I 400 bp overlap
ChIP RMG-I GSE104545.ARID1A.RMG-I 253 bp overlap
ARID1B 2 datasets
ChIP K-562 ENCSR822CCM.ARID1B.K-562 1016 bp overlap
ChIP K562 ENCFF938UXQ 521 bp overlap
ARID3A 1 dataset
ChIP HepG2 ENCFF341DES 525 bp overlap
ARNT 1 dataset
ChIP K562 ENCFF291CXK 425 bp overlap
ATF3 4 datasets
ChIP K-562 ENCSR028UIU.ATF3.K-562 616 bp overlap
ChIP K562 ENCFF604FPV 257 bp overlap
ChIP K562 ENCFF921JQW 665 bp overlap
ChIP K562 ENCFF921JQW 665 bp overlap
ATF7 3 datasets
ChIP K-562 ENCSR972ZBV.ATF7.K-562 324 bp overlap
ChIP K562 ENCFF308SKS 561 bp overlap
ChIP K562 ENCFF308SKS 561 bp overlap
ATOH7 4 datasets
Motif DE_36h DE_36h-ATOH7_MA1468.1 10 bp overlap
Motif DE_48h DE_48h-ATOH7_MA1468.1 10 bp overlap
Motif DE_60h DE_60h-ATOH7_MA1468.1 10 bp overlap
Motif DE_72h DE_72h-ATOH7_MA1468.1 10 bp overlap
Atoh1 4 datasets
Motif DE_36h DE_36h-Atoh1_MA0461.3 8 bp overlap
Motif DE_48h DE_48h-Atoh1_MA0461.3 8 bp overlap
Motif DE_60h DE_60h-Atoh1_MA0461.3 8 bp overlap
Motif DE_72h DE_72h-Atoh1_MA0461.3 8 bp overlap
BACH2 1 dataset
Motif DE_48h DE_48h-BACH2_MA1101.3 11 bp overlap
BHLHA15 4 datasets
Motif DE_36h DE_36h-BHLHA15_MA0607.2 10 bp overlap
Motif DE_48h DE_48h-BHLHA15_MA0607.2 10 bp overlap
Motif DE_60h DE_60h-BHLHA15_MA0607.2 10 bp overlap
Motif DE_72h DE_72h-BHLHA15_MA0607.2 10 bp overlap
BHLHE22 4 datasets
Motif DE_36h DE_36h-BHLHE22_MA0818.2 10 bp overlap
Motif DE_48h DE_48h-BHLHE22_MA0818.2 10 bp overlap
Motif DE_60h DE_60h-BHLHE22_MA0818.2 10 bp overlap
Motif DE_72h DE_72h-BHLHE22_MA0818.2 10 bp overlap
BHLHE23 4 datasets
Motif DE_36h DE_36h-BHLHE23_MA0817.2 10 bp overlap
Motif DE_48h DE_48h-BHLHE23_MA0817.2 10 bp overlap
Motif DE_60h DE_60h-BHLHE23_MA0817.2 10 bp overlap
Motif DE_72h DE_72h-BHLHE23_MA0817.2 10 bp overlap
BRD2 1 dataset
ChIP K-562_DMSO GSE138084.BRD2.K-562_DMSO 212 bp overlap
BRD3 1 dataset
ChIP K-562_iBET-BD1-IFNG GSE138084.BRD3.K-562_iBET-BD1-IFNG 204 bp overlap
BRD4 4 datasets
ChIP K-562_DMSO GSE138084.BRD4.K-562_DMSO 353 bp overlap
ChIP K-562_DMSO-IFNG GSE138084.BRD4.K-562_DMSO-IFNG 418 bp overlap
ChIP K-562_iBET-BD1-IFNG GSE138084.BRD4.K-562_iBET-BD1-IFNG 255 bp overlap
ChIP K-562_iBET-BD2-IFNG GSE138084.BRD4.K-562_iBET-BD2-IFNG 493 bp overlap
CBFA2T2 2 datasets
ChIP K-562 ENCSR699PVC.CBFA2T2.K-562 263 bp overlap
ChIP K562 ENCFF963TXY 292 bp overlap
CBFA2T3 3 datasets
ChIP K-562 ENCSR697YLJ.CBFA2T3.K-562 487 bp overlap
ChIP K-562 GSE142227.CBFA2T3.K-562 233 bp overlap
ChIP K562 ENCFF673OEZ 439 bp overlap
CC2D1A 1 dataset
ChIP K562 ENCFF567XUT 445 bp overlap
CHAMP1 2 datasets
ChIP K-562 ENCSR065XVO.CHAMP1.K-562 256 bp overlap
ChIP K562 ENCFF860ZIW 457 bp overlap
CREM 1 dataset
ChIP K562 ENCFF180STA 301 bp overlap
CTBP1 2 datasets
ChIP K-562 ENCSR201NQZ.CTBP1.K-562 232 bp overlap
ChIP K562 ENCFF403WPG 545 bp overlap
CUX2 3 datasets
Motif DE_48h DE_48h-CUX2_MA0755.2 9 bp overlap
Motif DE_60h DE_60h-CUX2_MA0755.2 9 bp overlap
Motif DE_72h DE_72h-CUX2_MA0755.2 9 bp overlap
CXXC5 1 dataset
ChIP K562 ENCFF497CZN 157 bp overlap
DACH1 1 dataset
ChIP K562 ENCFF574LOW 381 bp overlap
DPF2 5 datasets
ChIP K-562 ENCSR219BXP.DPF2.K-562 256 bp overlap
ChIP K-562 ENCSR219BXP.DPF2.K-562 473 bp overlap
ChIP K562 ENCFF739JDE 282 bp overlap
ChIP K562 ENCFF739JDE 497 bp overlap
ChIP K562 ENCFF775HUO 573 bp overlap
E2F5 1 dataset
ChIP K562 ENCFF688PUB 681 bp overlap
E4F1 3 datasets
ChIP K-562 ENCSR731LHZ.E4F1.K-562 258 bp overlap
ChIP K562 ENCFF622HMZ 601 bp overlap
ChIP K562 ENCFF622HMZ 601 bp overlap
EGR1 2 datasets
ChIP K562 ENCFF113OPQ 451 bp overlap
ChIP K562 ENCFF113OPQ 451 bp overlap
EHMT2 1 dataset
ChIP K-562 ENCSR175EOM.EHMT2.K-562 246 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 149 bp overlap
EP300 1 dataset
ChIP K562 ENCFF226VMS 317 bp overlap
EP400 2 datasets
ChIP K562 ENCFF850OZQ 745 bp overlap
ChIP K562 ENCFF850OZQ 745 bp overlap
ERF::NHLH1 4 datasets
Motif DE_36h DE_36h-ERFNHLH1_MA1938.2 16 bp overlap
Motif DE_48h DE_48h-ERFNHLH1_MA1938.2 16 bp overlap
Motif DE_60h DE_60h-ERFNHLH1_MA1938.2 16 bp overlap
Motif DE_72h DE_72h-ERFNHLH1_MA1938.2 16 bp overlap
ESRRA 4 datasets
Motif DE_36h DE_36h-ESRRA_MA0592.4 9 bp overlap
Motif DE_48h DE_48h-ESRRA_MA0592.4 9 bp overlap
Motif DE_60h DE_60h-ESRRA_MA0592.4 9 bp overlap
Motif DE_72h DE_72h-ESRRA_MA0592.4 9 bp overlap
ESRRB 4 datasets
Motif DE_36h DE_36h-ESRRB_MA0141.4 10 bp overlap
Motif DE_48h DE_48h-ESRRB_MA0141.4 10 bp overlap
Motif DE_60h DE_60h-ESRRB_MA0141.4 10 bp overlap
Motif DE_72h DE_72h-ESRRB_MA0141.4 10 bp overlap
ETV5::DRGX 4 datasets
Motif DE_36h DE_36h-ETV5DRGX_MA1944.2 12 bp overlap
Motif DE_48h DE_48h-ETV5DRGX_MA1944.2 12 bp overlap
Motif DE_60h DE_60h-ETV5DRGX_MA1944.2 12 bp overlap
Motif DE_72h DE_72h-ETV5DRGX_MA1944.2 12 bp overlap
ETV5::HOXA2 4 datasets
Motif DE_36h DE_36h-ETV5HOXA2_MA1948.2 12 bp overlap
Motif DE_48h DE_48h-ETV5HOXA2_MA1948.2 12 bp overlap
Motif DE_60h DE_60h-ETV5HOXA2_MA1948.2 12 bp overlap
Motif DE_72h DE_72h-ETV5HOXA2_MA1948.2 12 bp overlap
EZH2 1 dataset
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 198 bp overlap
FLI1::DRGX 4 datasets
Motif DE_36h DE_36h-FLI1DRGX_MA1949.2 14 bp overlap
Motif DE_48h DE_48h-FLI1DRGX_MA1949.2 14 bp overlap
Motif DE_60h DE_60h-FLI1DRGX_MA1949.2 14 bp overlap
Motif DE_72h DE_72h-FLI1DRGX_MA1949.2 14 bp overlap
FOSL1 5 datasets
ChIP K-562 ENCSR239ZLZ.FOSL1.K-562 520 bp overlap
ChIP K-562 ENCSR239ZLZ.FOSL1.K-562 294 bp overlap
ChIP K562 ENCFF455MKD 737 bp overlap
ChIP K562 ENCFF455MKD 737 bp overlap
ChIP K562 ENCFF455MKD 737 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 760 bp overlap
ChIP DE DE-FOXA2-2 727 bp overlap
ChIP pancreas_CARN1618 ERP008682.FOXA2.pancreas_CARN1618 218 bp overlap
FOXC1 3 datasets
Motif DE_48h DE_48h-FOXC1_MA0032.2 11 bp overlap
Motif DE_60h DE_60h-FOXC1_MA0032.2 11 bp overlap
Motif DE_72h DE_72h-FOXC1_MA0032.2 11 bp overlap
FOXK2 1 dataset
ChIP K562 ENCFF851PFH 457 bp overlap
FOXM1 1 dataset
ChIP K562 ENCFF255RHV 411 bp overlap
GABPB1 1 dataset
ChIP K562 ENCFF015GDS 551 bp overlap
GATA1 6 datasets
ChIP HUDEP-2_30min GSE104676.GATA1.HUDEP-2_30min 56 bp overlap
ChIP K-562 GSE107726.GATA1.K-562 282 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 260 bp overlap
ChIP K562 ENCFF094CMK 251 bp overlap
ChIP erythroblast ENCFF867JAR 605 bp overlap
ChIP erythroblast ENCSR000EXP.GATA1.erythroblast 480 bp overlap
GATA2 9 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
ChIP K-562 ENCSR000DKA.GATA2.K-562 442 bp overlap
ChIP K562 ENCFF088XQT 411 bp overlap
ChIP K562 ENCFF544PCK 251 bp overlap
ChIP K562 ENCFF830LLA 577 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 202 bp overlap
GATA4 8 datasets
ChIP DE DE-GATA4-1 775 bp overlap
ChIP DE DE-GATA4-2 1019 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 426 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 805 bp overlap
GATA5 4 datasets
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
Motif DE_48h DE_48h-GATA5_MA0766.3 8 bp overlap
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 13 datasets
ChIP DE DE-GATA6-1 784 bp overlap
ChIP DE DE-GATA6-2 894 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 298 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 507 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 593 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 371 bp overlap
ChIP foregut GSE117136.GATA6.foregut 369 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 333 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 262 bp overlap
GATAD2B 1 dataset
ChIP K562 ENCFF696VMK 401 bp overlap
Gata3 4 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HDAC1 3 datasets
ChIP K-562 ENCSR711VWL.HDAC1.K-562 283 bp overlap
ChIP K562 ENCFF928TKZ 227 bp overlap
ChIP K562 ENCFF968WBH 303 bp overlap
HDAC2 5 datasets
ChIP K-562 ENCSR893WSB.HDAC2.K-562 274 bp overlap
ChIP K-562 ENCSR893WSB.HDAC2.K-562 402 bp overlap
ChIP K-562 ENCSR893WSB.HDAC2.K-562 422 bp overlap
ChIP K562 ENCFF744ALD 359 bp overlap
ChIP K562 ENCFF919OMP 183 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR197ALX.HDGF.K-562 443 bp overlap
HLF 1 dataset
ChIP HepG2 ENCFF854JLR 245 bp overlap
HOXA9 4 datasets
Motif DE_36h DE_36h-HOXA9_MA0594.3 7 bp overlap
Motif DE_48h DE_48h-HOXA9_MA0594.3 7 bp overlap
Motif DE_60h DE_60h-HOXA9_MA0594.3 7 bp overlap
Motif DE_72h DE_72h-HOXA9_MA0594.3 7 bp overlap
Hand1::Tcf3 4 datasets
Motif DE_36h DE_36h-Hand1Tcf3_MA0092.2 9 bp overlap
Motif DE_48h DE_48h-Hand1Tcf3_MA0092.2 9 bp overlap
Motif DE_60h DE_60h-Hand1Tcf3_MA0092.2 9 bp overlap
Motif DE_72h DE_72h-Hand1Tcf3_MA0092.2 9 bp overlap
IKZF1 1 dataset
ChIP K-562 ENCSR395HWC.IKZF1.K-562 292 bp overlap
ILF3 1 dataset
ChIP K-562 GSE103215.ILF3.K-562 294 bp overlap
Irf1 4 datasets
Motif DE_36h DE_36h-Irf1_MA0050.4 11 bp overlap
Motif DE_48h DE_48h-Irf1_MA0050.4 11 bp overlap
Motif DE_60h DE_60h-Irf1_MA0050.4 11 bp overlap
Motif DE_72h DE_72h-Irf1_MA0050.4 11 bp overlap
JDP2 1 dataset
Motif DE_48h DE_48h-JDP2_MA0655.1 9 bp overlap
JUN 1 dataset
ChIP K-562 ENCSR000EFS.JUN.K-562 124 bp overlap
JUNB 1 dataset
ChIP K-562 ENCSR000DJY.JUNB.K-562 138 bp overlap
JUND 2 datasets
ChIP K-562 ENCSR000EGN.JUND.K-562 234 bp overlap
ChIP K562 ENCFF830LVJ 281 bp overlap
KDM1A 5 datasets
ChIP K-562 ENCSR360HRA.KDM1A.K-562 425 bp overlap
ChIP K-562 ENCSR908CMW.KDM1A.K-562 413 bp overlap
ChIP K562 ENCFF128TYE 332 bp overlap
ChIP K562 ENCFF133OLU 320 bp overlap
ChIP K562 ENCFF934ZRG 501 bp overlap
KLF16 1 dataset
ChIP K562 ENCFF464PIV 345 bp overlap
KLF5 4 datasets
Motif DE_36h DE_36h-KLF5_MA0599.1 10 bp overlap
Motif DE_48h DE_48h-KLF5_MA0599.1 10 bp overlap
Motif DE_60h DE_60h-KLF5_MA0599.1 10 bp overlap
Motif DE_72h DE_72h-KLF5_MA0599.1 10 bp overlap
L3MBTL2 2 datasets
ChIP K-562 ENCSR530XQI.L3MBTL2.K-562 418 bp overlap
ChIP K562 ENCFF320EQC 601 bp overlap
LDB1 2 datasets
ChIP K-562 GSE142227.LDB1.K-562 168 bp overlap
ChIP K-562 GSE142227.LDB1.K-562 324 bp overlap
LEF1 2 datasets
ChIP K562 ENCFF198WCP 457 bp overlap
ChIP K562 ENCFF198WCP 457 bp overlap
LIN54 4 datasets
Motif DE_36h DE_36h-LIN54_MA0619.2 7 bp overlap
Motif DE_48h DE_48h-LIN54_MA0619.2 7 bp overlap
Motif DE_60h DE_60h-LIN54_MA0619.2 7 bp overlap
Motif DE_72h DE_72h-LIN54_MA0619.2 7 bp overlap
MAX 2 datasets
ChIP K562 ENCFF398VJM 537 bp overlap
ChIP K562 ENCFF524IJO 118 bp overlap
MAZ 5 datasets
Motif DE_36h DE_36h-MAZ_MA1522.2 8 bp overlap
Motif DE_48h DE_48h-MAZ_MA1522.2 8 bp overlap
Motif DE_60h DE_60h-MAZ_MA1522.2 8 bp overlap
Motif DE_72h DE_72h-MAZ_MA1522.2 8 bp overlap
ChIP K562 ENCFF982GSZ 517 bp overlap
MCM7 1 dataset
ChIP K562 ENCFF081NIG 361 bp overlap
MEIS2 2 datasets
ChIP K-562 ENCSR851BNE.MEIS2.K-562 445 bp overlap
ChIP K562 ENCFF320GSD 318 bp overlap
MGA 1 dataset
ChIP K562 ENCFF140CEX 585 bp overlap
MITF 2 datasets
ChIP K-562 ENCSR797SWM.MITF.K-562 238 bp overlap
ChIP K562 ENCFF731XJJ 425 bp overlap
MLLT3 1 dataset
ChIP THP-1 GSE79899.MLLT3.THP-1 113 bp overlap
MNT 1 dataset
ChIP K562 ENCFF820IGH 651 bp overlap
MTA1 1 dataset
ChIP K562 ENCFF230ZKA 445 bp overlap
MTA2 2 datasets
ChIP K562 ENCFF441KCP 417 bp overlap
ChIP K562 ENCFF880VZB 141 bp overlap
MTA3 1 dataset
ChIP K562 ENCFF289UFB 537 bp overlap
MYOD1 1 dataset
ChIP RD_shSNAI2 GSE137168.MYOD1.RD_shSNAI2 181 bp overlap
NCOA1 2 datasets
ChIP K562 ENCFF395XLS 421 bp overlap
ChIP K562 ENCFF962VHQ 451 bp overlap
NCOR1 4 datasets
ChIP K-562 ENCSR798ILC.NCOR1.K-562 693 bp overlap
ChIP K-562 ENCSR910JAI.NCOR1.K-562 196 bp overlap
ChIP K562 ENCFF788MPU 263 bp overlap
ChIP K562 ENCFF866HRM 326 bp overlap
NEUROG1 4 datasets
Motif DE_36h DE_36h-NEUROG1_MA0623.2 10 bp overlap
Motif DE_48h DE_48h-NEUROG1_MA0623.2 10 bp overlap
Motif DE_60h DE_60h-NEUROG1_MA0623.2 10 bp overlap
Motif DE_72h DE_72h-NEUROG1_MA0623.2 10 bp overlap
NEUROG2 4 datasets
Motif DE_36h DE_36h-NEUROG2_MA0669.1 10 bp overlap
Motif DE_48h DE_48h-NEUROG2_MA0669.1 10 bp overlap
Motif DE_60h DE_60h-NEUROG2_MA0669.1 10 bp overlap
Motif DE_72h DE_72h-NEUROG2_MA0669.1 10 bp overlap
NFATC3 6 datasets
Motif DE_36h DE_36h-NFATC3_MA0625.3 6 bp overlap
Motif DE_48h DE_48h-NFATC3_MA0625.3 6 bp overlap
Motif DE_60h DE_60h-NFATC3_MA0625.3 6 bp overlap
Motif DE_72h DE_72h-NFATC3_MA0625.3 6 bp overlap
ChIP K-562 ENCSR051OUX.NFATC3.K-562 283 bp overlap
ChIP K562 ENCFF078EKB 451 bp overlap
NFE2 4 datasets
Motif DE_48h DE_48h-NFE2_MA0841.2 10 bp overlap
ChIP K-562 ENCSR000FCC.NFE2.K-562 104 bp overlap
ChIP K562 ENCFF047YKA 405 bp overlap
ChIP K562 ENCFF163BSI 265 bp overlap
NFIC 1 dataset
ChIP K562 ENCFF167YID 193 bp overlap
NOTCH3 1 dataset
ChIP TALL-1_GSI GSE104261.NOTCH3.TALL-1_GSI 335 bp overlap
NR2C1 5 datasets
Motif DE_36h DE_36h-NR2C1_MA1535.2 6 bp overlap
Motif DE_48h DE_48h-NR2C1_MA1535.2 6 bp overlap
Motif DE_60h DE_60h-NR2C1_MA1535.2 6 bp overlap
Motif DE_72h DE_72h-NR2C1_MA1535.2 6 bp overlap
ChIP K562 ENCFF568JLK 411 bp overlap
NR2C2 5 datasets
Motif DE_36h DE_36h-NR2C2_MA1536.2 6 bp overlap
Motif DE_48h DE_48h-NR2C2_MA1536.2 6 bp overlap
Motif DE_60h DE_60h-NR2C2_MA1536.2 6 bp overlap
Motif DE_72h DE_72h-NR2C2_MA1536.2 6 bp overlap
ChIP K562 ENCFF750AXF 911 bp overlap
NR2F1 6 datasets
Motif DE_36h DE_36h-NR2F1_MA0017.3 12 bp overlap
Motif DE_48h DE_48h-NR2F1_MA0017.3 12 bp overlap
Motif DE_60h DE_60h-NR2F1_MA0017.3 12 bp overlap
Motif DE_72h DE_72h-NR2F1_MA0017.3 12 bp overlap
ChIP K-562 ENCSR970NKQ.NR2F1.K-562 549 bp overlap
ChIP K562 ENCFF221HJH 449 bp overlap
NR2F2 3 datasets
ChIP K-562 ENCSR000BRS.NR2F2.K-562 364 bp overlap
ChIP K562 ENCFF004YPK 167 bp overlap
ChIP K562 ENCFF004YPK 391 bp overlap
NR2F6 2 datasets
ChIP K-562 ENCSR707QWA.NR2F6.K-562 333 bp overlap
ChIP K562 ENCFF674RQA 314 bp overlap
NR4A1 4 datasets
Motif DE_36h DE_36h-NR4A1_MA1112.3 8 bp overlap
Motif DE_48h DE_48h-NR4A1_MA1112.3 8 bp overlap
Motif DE_60h DE_60h-NR4A1_MA1112.3 8 bp overlap
Motif DE_72h DE_72h-NR4A1_MA1112.3 8 bp overlap
NR5A1 4 datasets
Motif DE_36h DE_36h-NR5A1_MA1540.3 12 bp overlap
Motif DE_48h DE_48h-NR5A1_MA1540.3 12 bp overlap
Motif DE_60h DE_60h-NR5A1_MA1540.3 12 bp overlap
Motif DE_72h DE_72h-NR5A1_MA1540.3 12 bp overlap
Nanog 4 datasets
Motif DE_36h DE_36h-Nanog_MA2339.1 7 bp overlap
Motif DE_48h DE_48h-Nanog_MA2339.1 7 bp overlap
Motif DE_60h DE_60h-Nanog_MA2339.1 7 bp overlap
Motif DE_72h DE_72h-Nanog_MA2339.1 7 bp overlap
Nfat5 1 dataset
Motif DE_48h DE_48h-Nfat5_MA0606.3 8 bp overlap
Nfatc1 4 datasets
Motif DE_36h DE_36h-Nfatc1_MA0624.3 6 bp overlap
Motif DE_48h DE_48h-Nfatc1_MA0624.3 6 bp overlap
Motif DE_60h DE_60h-Nfatc1_MA0624.3 6 bp overlap
Motif DE_72h DE_72h-Nfatc1_MA0624.3 6 bp overlap
Nfatc2 1 dataset
Motif DE_48h DE_48h-Nfatc2_MA0152.3 8 bp overlap
Nr1H2 4 datasets
Motif DE_36h DE_36h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_48h DE_48h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_60h DE_60h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_72h DE_72h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 4 datasets
Motif DE_36h DE_36h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_48h DE_48h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_60h DE_60h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_72h DE_72h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 4 datasets
Motif DE_36h DE_36h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_48h DE_48h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_60h DE_60h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_72h DE_72h-Nr1h3_MA2337.1 6 bp overlap
Nr2e1 4 datasets
Motif DE_36h DE_36h-Nr2e1_MA0676.1 9 bp overlap
Motif DE_48h DE_48h-Nr2e1_MA0676.1 9 bp overlap
Motif DE_60h DE_60h-Nr2e1_MA0676.1 9 bp overlap
Motif DE_72h DE_72h-Nr2e1_MA0676.1 9 bp overlap
Nr5A2 4 datasets
Motif DE_36h DE_36h-Nr5A2_MA0505.3 9 bp overlap
Motif DE_48h DE_48h-Nr5A2_MA0505.3 9 bp overlap
Motif DE_60h DE_60h-Nr5A2_MA0505.3 9 bp overlap
Motif DE_72h DE_72h-Nr5A2_MA0505.3 9 bp overlap
OLIG1 4 datasets
Motif DE_36h DE_36h-OLIG1_MA0826.1 10 bp overlap
Motif DE_48h DE_48h-OLIG1_MA0826.1 10 bp overlap
Motif DE_60h DE_60h-OLIG1_MA0826.1 10 bp overlap
Motif DE_72h DE_72h-OLIG1_MA0826.1 10 bp overlap
OLIG2 4 datasets
Motif DE_36h DE_36h-OLIG2_MA0678.1 10 bp overlap
Motif DE_48h DE_48h-OLIG2_MA0678.1 10 bp overlap
Motif DE_60h DE_60h-OLIG2_MA0678.1 10 bp overlap
Motif DE_72h DE_72h-OLIG2_MA0678.1 10 bp overlap
OLIG3 4 datasets
Motif DE_36h DE_36h-OLIG3_MA0827.1 10 bp overlap
Motif DE_48h DE_48h-OLIG3_MA0827.1 10 bp overlap
Motif DE_60h DE_60h-OLIG3_MA0827.1 10 bp overlap
Motif DE_72h DE_72h-OLIG3_MA0827.1 10 bp overlap
ONECUT2 3 datasets
Motif DE_48h DE_48h-ONECUT2_MA0756.3 8 bp overlap
Motif DE_60h DE_60h-ONECUT2_MA0756.3 8 bp overlap
Motif DE_72h DE_72h-ONECUT2_MA0756.3 8 bp overlap
PAX3-FOXO1 1 dataset
ChIP Hs-352-Sk_PAX3-FOXO1-vector GSE83725.PAX3-FOXO1.Hs-352-Sk_PAX3-FOXO1-vector 177 bp overlap
PBX1 4 datasets
Motif DE_36h DE_36h-PBX1_MA0070.2 9 bp overlap
Motif DE_48h DE_48h-PBX1_MA0070.2 9 bp overlap
Motif DE_60h DE_60h-PBX1_MA0070.2 9 bp overlap
Motif DE_72h DE_72h-PBX1_MA0070.2 9 bp overlap
PBX2 2 datasets
ChIP K-562 ENCSR263DFP.PBX2.K-562 554 bp overlap
ChIP K562 ENCFF286KMN 222 bp overlap
PHF8 1 dataset
ChIP Hep-G2 ENCSR604VAE.PHF8.Hep-G2 177 bp overlap
PKNOX1 3 datasets
ChIP GM12878 ENCFF589FCY 437 bp overlap
ChIP K-562 ENCSR115SMW.PKNOX1.K-562 233 bp overlap
ChIP K562 ENCFF236IUS 422 bp overlap
POU2F3 4 datasets
Motif DE_36h DE_36h-POU2F3_MA0627.3 9 bp overlap
Motif DE_48h DE_48h-POU2F3_MA0627.3 9 bp overlap
Motif DE_60h DE_60h-POU2F3_MA0627.3 9 bp overlap
Motif DE_72h DE_72h-POU2F3_MA0627.3 9 bp overlap
POU3F1 8 datasets
Motif DE_36h DE_36h-POU3F1_MA0786.2 10 bp overlap
Motif DE_36h DE_36h-POU3F1_MA0786.2 10 bp overlap
Motif DE_48h DE_48h-POU3F1_MA0786.2 10 bp overlap
Motif DE_48h DE_48h-POU3F1_MA0786.2 10 bp overlap
Motif DE_60h DE_60h-POU3F1_MA0786.2 10 bp overlap
Motif DE_60h DE_60h-POU3F1_MA0786.2 10 bp overlap
Motif DE_72h DE_72h-POU3F1_MA0786.2 10 bp overlap
Motif DE_72h DE_72h-POU3F1_MA0786.2 10 bp overlap
POU4F1 4 datasets
Motif DE_36h DE_36h-POU4F1_MA0790.2 12 bp overlap
Motif DE_48h DE_48h-POU4F1_MA0790.2 12 bp overlap
Motif DE_60h DE_60h-POU4F1_MA0790.2 12 bp overlap
Motif DE_72h DE_72h-POU4F1_MA0790.2 12 bp overlap
POU4F2 4 datasets
Motif DE_36h DE_36h-POU4F2_MA0683.2 15 bp overlap
Motif DE_48h DE_48h-POU4F2_MA0683.2 15 bp overlap
Motif DE_60h DE_60h-POU4F2_MA0683.2 15 bp overlap
Motif DE_72h DE_72h-POU4F2_MA0683.2 15 bp overlap
POU4F3 4 datasets
Motif DE_36h DE_36h-POU4F3_MA0791.2 12 bp overlap
Motif DE_48h DE_48h-POU4F3_MA0791.2 12 bp overlap
Motif DE_60h DE_60h-POU4F3_MA0791.2 12 bp overlap
Motif DE_72h DE_72h-POU4F3_MA0791.2 12 bp overlap
PRDM10 3 datasets
ChIP HepG2 ENCFF324FNA 521 bp overlap
ChIP K-562 ENCSR120MPG.PRDM10.K-562 598 bp overlap
ChIP K562 ENCFF740YLK 412 bp overlap
PROX1 2 datasets
ChIP HepG2 ENCFF016ZJS 481 bp overlap
ChIP HepG2 ENCFF016ZJS 481 bp overlap
Pou5f1::Sox2 4 datasets
Motif DE_36h DE_36h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_48h DE_48h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_60h DE_60h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_72h DE_72h-Pou5f1Sox2_MA0142.1 15 bp overlap
Pparg::Rxra 4 datasets
Motif DE_36h DE_36h-PpargRxra_MA0065.3 13 bp overlap
Motif DE_48h DE_48h-PpargRxra_MA0065.3 13 bp overlap
Motif DE_60h DE_60h-PpargRxra_MA0065.3 13 bp overlap
Motif DE_72h DE_72h-PpargRxra_MA0065.3 13 bp overlap
Ptf1A 4 datasets
Motif DE_36h DE_36h-Ptf1A_MA1618.2 9 bp overlap
Motif DE_48h DE_48h-Ptf1A_MA1618.2 9 bp overlap
Motif DE_60h DE_60h-Ptf1A_MA1618.2 9 bp overlap
Motif DE_72h DE_72h-Ptf1A_MA1618.2 9 bp overlap
RAD21 1 dataset
ChIP K-562_RF2-insertion GSE140868.RAD21.K-562_RF2-insertion 118 bp overlap
RBM39 1 dataset
ChIP K-562 ENCSR764OXF.RBM39.K-562 149 bp overlap
RCOR1 2 datasets
ChIP K562 ENCFF216EEJ 297 bp overlap
ChIP K562 ENCFF721RTS 345 bp overlap
REL 4 datasets
Motif DE_36h DE_36h-REL_MA0101.1 10 bp overlap
Motif DE_48h DE_48h-REL_MA0101.1 10 bp overlap
Motif DE_60h DE_60h-REL_MA0101.1 10 bp overlap
Motif DE_72h DE_72h-REL_MA0101.1 10 bp overlap
RFX1 1 dataset
ChIP K562 ENCFF421AVO 465 bp overlap
RNF2 2 datasets
ChIP K562 ENCFF061ATI 445 bp overlap
ChIP K562 ENCFF653BQJ 286 bp overlap
SETDB1 1 dataset
ChIP WN8532 GSE36579.SETDB1.WN8532 207 bp overlap
SIN3A 1 dataset
ChIP K-562 ENCSR920BLG.SIN3A.K-562 193 bp overlap
SIX2 1 dataset
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 93 bp overlap
SMAD2 4 datasets
Motif DE_36h DE_36h-SMAD2_MA1964.2 6 bp overlap
Motif DE_48h DE_48h-SMAD2_MA1964.2 6 bp overlap
Motif DE_60h DE_60h-SMAD2_MA1964.2 6 bp overlap
Motif DE_72h DE_72h-SMAD2_MA1964.2 6 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 484 bp overlap
SMAD2_3 2 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 471 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 346 bp overlap
SMAD4 2 datasets
ChIP K562 ENCFF628RBP 541 bp overlap
ChIP K562 ENCFF628RBP 541 bp overlap
SMARCA4 5 datasets
ChIP K-562 ENCSR587OQL.SMARCA4.K-562 427 bp overlap
ChIP K-562 ENCSR643VTW.SMARCA4.K-562 287 bp overlap
ChIP K-562 ENCSR643VTW.SMARCA4.K-562 324 bp overlap
ChIP K562 ENCFF316MCJ 485 bp overlap
ChIP K562 ENCFF506JCB 403 bp overlap
SMARCC2 1 dataset
ChIP K562 ENCFF368GSR 497 bp overlap
SMARCE1 3 datasets
ChIP K-562 ENCSR157TCS.SMARCE1.K-562 414 bp overlap
ChIP K-562 ENCSR157TCS.SMARCE1.K-562 274 bp overlap
ChIP K562 ENCFF690CFF 304 bp overlap
SOX13 5 datasets
Motif DE_36h DE_36h-SOX13_MA1120.2 7 bp overlap
Motif DE_48h DE_48h-SOX13_MA1120.2 7 bp overlap
Motif DE_60h DE_60h-SOX13_MA1120.2 7 bp overlap
Motif DE_72h DE_72h-SOX13_MA1120.2 7 bp overlap
ChIP HepG2 ENCFF062VSQ 138 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 464 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 654 bp overlap
SOX2 4 datasets
Motif DE_36h DE_36h-SOX2_MA0143.5 7 bp overlap
Motif DE_48h DE_48h-SOX2_MA0143.5 7 bp overlap
Motif DE_60h DE_60h-SOX2_MA0143.5 7 bp overlap
Motif DE_72h DE_72h-SOX2_MA0143.5 7 bp overlap
SOX6 2 datasets
ChIP K-562 ENCSR788RSW.SOX6.K-562 554 bp overlap
ChIP K562 ENCFF059YCJ 472 bp overlap
SP4 4 datasets
Motif DE_36h DE_36h-SP4_MA0685.2 9 bp overlap
Motif DE_48h DE_48h-SP4_MA0685.2 9 bp overlap
Motif DE_60h DE_60h-SP4_MA0685.2 9 bp overlap
Motif DE_72h DE_72h-SP4_MA0685.2 9 bp overlap
SP5 4 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
Motif DE_72h DE_72h-SP5_MA1965.2 6 bp overlap
SRF 1 dataset
ChIP GM12878 ENCFF878IIX 397 bp overlap
STAT5A 1 dataset
ChIP K562 ENCFF226BTJ 341 bp overlap
Sox17 4 datasets
Motif DE_36h DE_36h-Sox17_MA0078.3 10 bp overlap
Motif DE_48h DE_48h-Sox17_MA0078.3 10 bp overlap
Motif DE_60h DE_60h-Sox17_MA0078.3 10 bp overlap
Motif DE_72h DE_72h-Sox17_MA0078.3 10 bp overlap
Sox3 4 datasets
Motif DE_36h DE_36h-Sox3_MA0514.3 7 bp overlap
Motif DE_48h DE_48h-Sox3_MA0514.3 7 bp overlap
Motif DE_60h DE_60h-Sox3_MA0514.3 7 bp overlap
Motif DE_72h DE_72h-Sox3_MA0514.3 7 bp overlap
Sox5 4 datasets
Motif DE_36h DE_36h-Sox5_MA0087.3 8 bp overlap
Motif DE_48h DE_48h-Sox5_MA0087.3 8 bp overlap
Motif DE_60h DE_60h-Sox5_MA0087.3 8 bp overlap
Motif DE_72h DE_72h-Sox5_MA0087.3 8 bp overlap
Sox6 4 datasets
Motif DE_36h DE_36h-Sox6_MA0515.1 10 bp overlap
Motif DE_48h DE_48h-Sox6_MA0515.1 10 bp overlap
Motif DE_60h DE_60h-Sox6_MA0515.1 10 bp overlap
Motif DE_72h DE_72h-Sox6_MA0515.1 10 bp overlap
Sox7 4 datasets
Motif DE_36h DE_36h-Sox7_MA2095.1 10 bp overlap
Motif DE_48h DE_48h-Sox7_MA2095.1 10 bp overlap
Motif DE_60h DE_60h-Sox7_MA2095.1 10 bp overlap
Motif DE_72h DE_72h-Sox7_MA2095.1 10 bp overlap
TAF9B 1 dataset
ChIP K562 ENCFF121ZIF 489 bp overlap
TAL1 10 datasets
ChIP K-562 ENCSR000EHB.TAL1.K-562 313 bp overlap
ChIP K-562 ENCSR106FRG.TAL1.K-562 256 bp overlap
ChIP K-562 GSE107726.TAL1.K-562 193 bp overlap
ChIP K-562_dCas9-LSD1 GSE132212.TAL1.K-562_dCas9-LSD1 155 bp overlap
ChIP K-562_enCRISPRi-KL GSE132212.TAL1.K-562_enCRISPRi-KL 296 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.TAL1.K-562_enCRISPRi-LK 299 bp overlap
ChIP K-562_sgGal4 GSE132212.TAL1.K-562_sgGal4 166 bp overlap
ChIP K562 ENCFF620GMX 352 bp overlap
ChIP K562 ENCFF661CCK 277 bp overlap
ChIP ProEs GSE59087.TAL1.ProEs 151 bp overlap
TAL1::TCF3 4 datasets
Motif DE_36h DE_36h-TAL1TCF3_MA0091.2 10 bp overlap
Motif DE_48h DE_48h-TAL1TCF3_MA0091.2 10 bp overlap
Motif DE_60h DE_60h-TAL1TCF3_MA0091.2 10 bp overlap
Motif DE_72h DE_72h-TAL1TCF3_MA0091.2 10 bp overlap
TBP 1 dataset
ChIP K-562 GSE55306.TBP.K-562 188 bp overlap
TBX21 1 dataset
Motif DE_48h DE_48h-TBX21_MA0690.3 10 bp overlap
TCF12 4 datasets
ChIP K-562 ENCSR744WOO.TCF12.K-562 372 bp overlap
ChIP K-562 ENCSR189TRZ.TCF12.K-562 211 bp overlap
ChIP K562 ENCFF909RDY 397 bp overlap
ChIP K562 ENCFF931DJY 326 bp overlap
TCF3 2 datasets
ChIP K-562 ENCSR970OJY.TCF3.K-562 366 bp overlap
ChIP K562 ENCFF319QZT 166 bp overlap
TEAD1 1 dataset
ChIP K562 ENCFF465AQA 297 bp overlap
TEAD4 1 dataset
ChIP K562 ENCFF673NIK 242 bp overlap
TFAP4 6 datasets
Motif DE_36h DE_36h-TFAP4_MA1570.1 10 bp overlap
Motif DE_48h DE_48h-TFAP4_MA1570.1 10 bp overlap
Motif DE_60h DE_60h-TFAP4_MA1570.1 10 bp overlap
Motif DE_72h DE_72h-TFAP4_MA1570.1 10 bp overlap
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 159 bp overlap
ChIP K562 ENCFF727PXG 531 bp overlap
TFDP1 4 datasets
Motif DE_36h DE_36h-TFDP1_MA1122.2 8 bp overlap
Motif DE_48h DE_48h-TFDP1_MA1122.2 8 bp overlap
Motif DE_60h DE_60h-TFDP1_MA1122.2 8 bp overlap
Motif DE_72h DE_72h-TFDP1_MA1122.2 8 bp overlap
TRIM24 3 datasets
ChIP K-562 ENCSR907MZR.TRIM24.K-562 242 bp overlap
ChIP K562 ENCFF284DKY 381 bp overlap
ChIP K562 ENCFF616RIL 451 bp overlap
TRIM28 2 datasets
ChIP WIBR3_NAIVE GSE84382.TRIM28.WIBR3_NAIVE 411 bp overlap
ChIP WIBR3_NAIVE GSE84382.TRIM28.WIBR3_NAIVE 138 bp overlap
TRPS1 4 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
Tbx6 1 dataset
Motif DE_48h DE_48h-Tbx6_MA1567.3 9 bp overlap
ZBTB17 1 dataset
ChIP K562 ENCFF731UTU 445 bp overlap
ZBTB2 1 dataset
ChIP K562 ENCFF290ESQ 431 bp overlap
ZBTB7A 1 dataset
ChIP HUDEP-2 GSE103445.ZBTB7A.HUDEP-2 337 bp overlap
ZEB2 3 datasets
ChIP K-562 ENCSR004GKA.ZEB2.K-562 908 bp overlap
ChIP K562 ENCFF795CMH 252 bp overlap
ChIP K562 ENCFF975RXS 461 bp overlap
ZIM3 4 datasets
Motif DE_36h DE_36h-ZIM3_MA1709.2 11 bp overlap
Motif DE_48h DE_48h-ZIM3_MA1709.2 11 bp overlap
Motif DE_60h DE_60h-ZIM3_MA1709.2 11 bp overlap
Motif DE_72h DE_72h-ZIM3_MA1709.2 11 bp overlap
ZNF148 4 datasets
Motif DE_36h DE_36h-ZNF148_MA1653.2 10 bp overlap
Motif DE_48h DE_48h-ZNF148_MA1653.2 10 bp overlap
Motif DE_60h DE_60h-ZNF148_MA1653.2 10 bp overlap
Motif DE_72h DE_72h-ZNF148_MA1653.2 10 bp overlap
ZNF24 2 datasets
ChIP K-562 ENCSR695EQB.ZNF24.K-562 371 bp overlap
ChIP K562 ENCFF781QQQ 361 bp overlap
ZNF418 1 dataset
Motif DE_48h DE_48h-ZNF418_MA1980.1 15 bp overlap
ZNF479 1 dataset
ChIP HEK293T GSE78099.ZNF479.HEK293T 82 bp overlap
ZNF524 3 datasets
Motif DE_48h DE_48h-ZNF524_MA2096.1 9 bp overlap
Motif DE_60h DE_60h-ZNF524_MA2096.1 9 bp overlap
Motif DE_72h DE_72h-ZNF524_MA2096.1 9 bp overlap
ZNF528 4 datasets
Motif DE_36h DE_36h-ZNF528_MA1597.1 17 bp overlap
Motif DE_48h DE_48h-ZNF528_MA1597.1 17 bp overlap
Motif DE_60h DE_60h-ZNF528_MA1597.1 17 bp overlap
Motif DE_72h DE_72h-ZNF528_MA1597.1 17 bp overlap
ZNF592 2 datasets
ChIP K562 ENCFF547OSS 605 bp overlap
ChIP K562 ENCFF547OSS 605 bp overlap
ZNF639 1 dataset
ChIP K562 ENCFF271FQR 741 bp overlap
ZNF816 4 datasets
Motif DE_36h DE_36h-ZNF816_MA1719.2 15 bp overlap
Motif DE_48h DE_48h-ZNF816_MA1719.2 15 bp overlap
Motif DE_60h DE_60h-ZNF816_MA1719.2 15 bp overlap
Motif DE_72h DE_72h-ZNF816_MA1719.2 15 bp overlap