chr12 : 52,947,826 – 52,948,366
540 bp 114 TFs 2 linked genes
This 540 bp open chromatin element is linked to KRT18 and KRT8 and is bound by 114 transcription factors.
Linked Genes
2 genes
Distance
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
KRT18 750 bp At TSS Proximity
KRT8 1.5 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr12:52,942,826 – 52,953,366
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
114 transcription factors
Source
Cell type
AFF4 1 dataset
ChIP MCF-7 GSE144036.AFF4.MCF-7 141 bp overlap
AR 1 dataset
ChIP LNCaP_FOXA1 ERP003503.AR.LNCaP_FOXA1 143 bp overlap
ARID1A 1 dataset
ChIP MCF-7_4-hydroxytamoxifen GSE123284.ARID1A.MCF-7_4-hydroxytamoxifen 352 bp overlap
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 343 bp overlap
BCL6 1 dataset
ChIP CD4 GSE59933.BCL6.CD4 143 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 333 bp overlap
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 219 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 540 bp overlap
BRD1 2 datasets
ChIP HUES-64 GSE104059.BRD1.HUES-64 349 bp overlap
ChIP RKO GSE47190.BRD1.RKO 156 bp overlap
BRD4 7 datasets
ChIP COLO-205 GSE73319.BRD4.COLO-205 540 bp overlap
ChIP HCT-15 GSE73319.BRD4.HCT-15 540 bp overlap
ChIP MCF-7_ARID1A-KO_4-OHT GSE123284.BRD4.MCF-7_ARID1A-KO_4-OHT 540 bp overlap
ChIP MCF-7_parental GSE123284.BRD4.MCF-7_parental 540 bp overlap
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.BRD4.MCF-7_parental_4-hydroxytamoxifen 540 bp overlap
ChIP PC-3_GDC-R_SAHA GSE137207.BRD4.PC-3_GDC-R_SAHA 280 bp overlap
ChIP SW480 GSE73319.BRD4.SW480 540 bp overlap
CAMTA2 1 dataset
ChIP HepG2 ENCFF305ZLM 241 bp overlap
CBFB 1 dataset
ChIP Hep-G2 ENCSR479RZV.CBFB.Hep-G2 92 bp overlap
CHD1 3 datasets
ChIP HeLa-S3 ENCSR392EAA.CHD1.HeLa-S3 147 bp overlap
ChIP MCF-7 ENCSR360JOC.CHD1.MCF-7 540 bp overlap
ChIP WA01 ENCSR000AQK.CHD1.WA01 507 bp overlap
CHD4 1 dataset
ChIP A-549 ENCSR550SCU.CHD4.A-549 416 bp overlap
CHD7 1 dataset
ChIP WA01 ENCSR000AVA.CHD7.WA01 193 bp overlap
CREB1 1 dataset
ChIP WA01 ENCSR000BSN.CREB1.WA01 120 bp overlap
CREBBP 2 datasets
ChIP PC-3 GSE147455.CREBBP.PC-3 116 bp overlap
ChIP PC-3_GDC-resist GSE147455.CREBBP.PC-3_GDC-resist 255 bp overlap
CRY1 1 dataset
ChIP U2OS GSE44236.CRY1.U2OS 251 bp overlap
DDX5 1 dataset
ChIP NTERA2 GSE58641.DDX5.NTERA2 143 bp overlap
E2F1 1 dataset
ChIP MDA-MB-231 GSE95303.E2F1.MDA-MB-231 231 bp overlap
E2F4 1 dataset
ChIP K-562 ENCSR000EWL.E2F4.K-562 128 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 155 bp overlap
EMX1 1 dataset
ChIP WTC11 ENCFF692RZJ 540 bp overlap
EP300 1 dataset
ChIP HeLa-S3 ENCSR680OFU.EP300.HeLa-S3 189 bp overlap
ESR1 7 datasets
ChIP MCF-7_DHT GSE99626.ESR1.MCF-7_DHT 428 bp overlap
ChIP T-47D GSE148277.ESR1.T-47D 232 bp overlap
ChIP T-47D_DMSO GSE148277.ESR1.T-47D_DMSO 344 bp overlap
ChIP T-47D_Y537S GSE148277.ESR1.T-47D_Y537S 412 bp overlap
ChIP T-47D_Y537S_E2 GSE148277.ESR1.T-47D_Y537S_E2 540 bp overlap
ChIP breast_tumor_Male_20 GSE104399.ESR1.breast_tumor_Male_20 172 bp overlap
ChIP breast_tumor_Male_21 GSE104399.ESR1.breast_tumor_Male_21 330 bp overlap
ESRRA 1 dataset
ChIP SK-BR-3_HRG GSE81651.ESRRA.SK-BR-3_HRG 282 bp overlap
ETS1 3 datasets
ChIP HEY-A8 GSE101832.ETS1.HEY-A8 422 bp overlap
ChIP HUVEC-C_VEGF_12H GSE41166.ETS1.HUVEC-C_VEGF_12H 190 bp overlap
ChIP cardiomyocyte_D9 GSE129986.ETS1.cardiomyocyte_D9 165 bp overlap
EZH2 1 dataset
ChIP THP-1 GSE135024.EZH2.THP-1 192 bp overlap
FOS 1 dataset
ChIP HCT-116_A7A GSE152144.FOS.HCT-116_A7A 420 bp overlap
FOSB 1 dataset
ChIP A549 ENCFF029EEU 341 bp overlap
FOXA1 1 dataset
ChIP T-47D_shRNF2_8h GSE137579.FOXA1.T-47D_shRNF2_8h 106 bp overlap
GATA6 3 datasets
ChIP DE_D1 S09-DE-d1-GATA6-exp1 270 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 310 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 334 bp overlap
GTF3C2 1 dataset
ChIP H9 GSE94418.GTF3C2.H9 144 bp overlap
HDAC2 1 dataset
ChIP PC-3 GSE147455.HDAC2.PC-3 329 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR563YDA.HDGF.K-562 355 bp overlap
HOXB13 2 datasets
ChIP A-549 ENCSR967ZMR.HOXB13.A-549 255 bp overlap
ChIP A549 ENCFF870NOA 351 bp overlap
JUN 9 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 349 bp overlap
ChIP DE_D1 S13-DE-d1-JUN-exp1 284 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 540 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 261 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 493 bp overlap
ChIP HUES-8 GSE109524.JUN.HUES-8 472 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 480 bp overlap
ChIP MDA-BoM-1833 GSE112444.JUN.MDA-BoM-1833 305 bp overlap
ChIP MDA-MB-231 GSE112444.JUN.MDA-MB-231 262 bp overlap
KAT7 1 dataset
ChIP K562 ENCFF175ZTN 476 bp overlap
KDM1A 1 dataset
ChIP NCI-H526_DMSO GSE66297.KDM1A.NCI-H526_DMSO 141 bp overlap
KMT2A 1 dataset
ChIP Hep-G2 ENCSR692GFR.KMT2A.Hep-G2 231 bp overlap
MBD3 1 dataset
ChIP MCF-7 GSE44737.MBD3.MCF-7 516 bp overlap
MED1 4 datasets
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 224 bp overlap
ChIP HUVEC-C_siCTL GSE124892.MED1.HUVEC-C_siCTL 227 bp overlap
ChIP HUVEC-C_siERG GSE124892.MED1.HUVEC-C_siERG 314 bp overlap
ChIP cardiomyocyte_5 GSE85628.MED1.cardiomyocyte_5 202 bp overlap
MSX2 2 datasets
ChIP MCF-7 ENCFF179YRV 297 bp overlap
ChIP MCF-7 ENCFF179YRV 244 bp overlap
MYBL2 1 dataset
ChIP A-673 GSE119971.MYBL2.A-673 129 bp overlap
NANOG 5 datasets
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 126 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 427 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 247 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 435 bp overlap
ChIP hESC GSE18292.NANOG.hESC 93 bp overlap
NCAPH2 2 datasets
ChIP RMG-I_ARID1A-KO GSE120058.NCAPH2.RMG-I_ARID1A-KO 190 bp overlap
ChIP RMG-I_ARID1A-KO GSE120058.NCAPH2.RMG-I_ARID1A-KO 158 bp overlap
NKX2-1 1 dataset
ChIP NCI-H3122 GSE39998.NKX2-1.NCI-H3122 200 bp overlap
NR3C1 2 datasets
ChIP K562 ENCFF877YZJ 91 bp overlap
ChIP T47D-A1-2_Dex GSE112491.NR3C1.T47D-A1-2_Dex 87 bp overlap
OTX1 1 dataset
ChIP MCF-7 ENCFF645GYL 317 bp overlap
PAF1 1 dataset
ChIP HCT-116_ab20662 GSE97527.PAF1.HCT-116_ab20662 264 bp overlap
PCBP1 2 datasets
ChIP Hep-G2 ENCSR872EVQ.PCBP1.Hep-G2 189 bp overlap
ChIP Hep-G2 GSE120104.PCBP1.Hep-G2 189 bp overlap
PCBP2 2 datasets
ChIP Hep-G2 ENCSR945NSF.PCBP2.Hep-G2 234 bp overlap
ChIP Hep-G2 GSE120104.PCBP2.Hep-G2 207 bp overlap
PGR 4 datasets
ChIP MCF-7_PROG GSE68355.PGR.MCF-7_PROG 224 bp overlap
ChIP MCF-7_R5020 GSE68355.PGR.MCF-7_R5020 212 bp overlap
ChIP T-47D GSE126859.PGR.T-47D 168 bp overlap
ChIP T-47D_R5020-A1 GSE126859.PGR.T-47D_R5020-A1 194 bp overlap
PHIP 4 datasets
ChIP HCT-116_MLL1-KO__ab833 GSE101646.PHIP.HCT-116_MLL1-KO__ab833 540 bp overlap
ChIP HCT-116_MLL4-SET-KO_ab833 GSE101646.PHIP.HCT-116_MLL4-SET-KO_ab833 249 bp overlap
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 540 bp overlap
ChIP HCT-116_ab834 GSE101646.PHIP.HCT-116_ab834 540 bp overlap
POLR2A 8 datasets
ChIP GM23338 ENCFF450WCS 456 bp overlap
ChIP H1 ENCFF566JSR 310 bp overlap
ChIP H1 ENCFF833NJP 316 bp overlap
ChIP HepG2 ENCFF736SLT 322 bp overlap
ChIP MCF-7 ENCFF309IKZ 255 bp overlap
ChIP MCF-7 ENCFF411WCU 280 bp overlap
ChIP Panc1 ENCFF290KAB 391 bp overlap
ChIP Panc1 ENCFF290KAB 187 bp overlap
POU2F1 1 dataset
ChIP T-47D GSE148277.POU2F1.T-47D 341 bp overlap
POU2F3 1 dataset
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 186 bp overlap
POU5F1 5 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 540 bp overlap
ChIP GM23338 ENCFF333SNB 311 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 430 bp overlap
ChIP hESC ENCSR264RJX.POU5F1.hESC 158 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 150 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 504 bp overlap
PRPF4 1 dataset
ChIP Hep-G2 GSE120104.PRPF4.Hep-G2 152 bp overlap
RAD21 1 dataset
ChIP HCT-116 GSE131606.RAD21.HCT-116 398 bp overlap
RARA 1 dataset
ChIP U-937_ATRA-treated_RAR GSE98006.RARA.U-937_ATRA-treated_RAR 110 bp overlap
RBFOX2 2 datasets
ChIP K-562 GSE120104.RBFOX2.K-562 430 bp overlap
ChIP K-562 ENCSR822LBD.RBFOX2.K-562 384 bp overlap
RBM39 3 datasets
ChIP Hep-G2 ENCSR339JTP.RBM39.Hep-G2 164 bp overlap
ChIP Hep-G2 GSE120104.RBM39.Hep-G2 158 bp overlap
ChIP K-562 ENCSR764OXF.RBM39.K-562 53 bp overlap
SIN3A 2 datasets
ChIP A-549 ENCSR513XQX.SIN3A.A-549 323 bp overlap
ChIP WA01 ENCSR000EBO.SIN3A.WA01 63 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 284 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 455 bp overlap
SMAD3 2 datasets
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 94 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 168 bp overlap
SMAD4 1 dataset
ChIP K562 ENCFF316DFN 371 bp overlap
SMAD5 2 datasets
ChIP K-562 ENCSR000FCD.SMAD5.K-562 122 bp overlap
ChIP K-562 ENCSR000FCD.SMAD5.K-562 132 bp overlap
SMAD7 2 datasets
ChIP HepG2 ENCFF850FXR 424 bp overlap
ChIP HepG2 ENCFF850FXR 540 bp overlap
SMARCA4 5 datasets
ChIP MCF-7 GSE140185.SMARCA4.MCF-7 274 bp overlap
ChIP MCF-7_ARID1A-KO_clone14 GSE123284.SMARCA4.MCF-7_ARID1A-KO_clone14 298 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 540 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 225 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 392 bp overlap
SMARCB1 3 datasets
ChIP HeLa-S3 ENCFF733PLR 367 bp overlap
ChIP hiPSC GSE124903.SMARCB1.hiPSC 190 bp overlap
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 169 bp overlap
SMARCC1 5 datasets
ChIP DE_D1 S10-DE-d1-BAF155-exp1 257 bp overlap
ChIP DE_D1 S15-DE-d1-BAF155-exp1 442 bp overlap
ChIP ESC S25-ESC-d0-BAF155-exp1 534 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 249 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 181 bp overlap
SMC1 1 dataset
ChIP DKO GSE131606.SMC1.DKO 264 bp overlap
SMC3 1 dataset
ChIP A549 ENCFF747SCJ 231 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 193 bp overlap
SP1 1 dataset
ChIP Hep-G2 ENCSR460YAM.SP1.Hep-G2 113 bp overlap
SP4 1 dataset
ChIP H1 ENCFF473YOB 157 bp overlap
SRSF3 2 datasets
ChIP K-562 GSE120104.SRSF3.K-562 333 bp overlap
ChIP K-562 ENCSR268QIQ.SRSF3.K-562 333 bp overlap
SSRP1 2 datasets
ChIP HepG2 ENCFF540BLL 440 bp overlap
ChIP HepG2 ENCFF540BLL 432 bp overlap
STAT6 1 dataset
ChIP HepG2 ENCFF370LZV 216 bp overlap
SUPT5H 1 dataset
ChIP DLD-1_NELFCD-AID GSE144786.SUPT5H.DLD-1_NELFCD-AID 540 bp overlap
SUPT6H 1 dataset
ChIP HCT-116 GSE130509.SUPT6H.HCT-116 345 bp overlap
T 2 datasets
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 185 bp overlap
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 201 bp overlap
TAF1 1 dataset
ChIP H1 ENCFF478SZO 140 bp overlap
TARDBP 3 datasets
ChIP K-562 GSE120104.TARDBP.K-562 258 bp overlap
ChIP K-562 ENCSR429XTR.TARDBP.K-562 195 bp overlap
ChIP K562 ENCFF021QCU 345 bp overlap
TBP 5 datasets
ChIP WA01 ENCSR000ECB.TBP.WA01 149 bp overlap
ChIP hESC GSE122298.TBP.hESC 363 bp overlap
ChIP hESC_10h GSE122298.TBP.hESC_10h 305 bp overlap
ChIP hESC_2h GSE122298.TBP.hESC_2h 313 bp overlap
ChIP hESC_8h GSE122298.TBP.hESC_8h 110 bp overlap
TBX5 5 datasets
ChIP G296S GSE85628.TBX5.G296S 171 bp overlap
ChIP G296S_2 GSE85628.TBX5.G296S_2 171 bp overlap
ChIP G296S_4 GSE85628.TBX5.G296S_4 406 bp overlap
ChIP cardiomyocyte GSE85628.TBX5.cardiomyocyte 316 bp overlap
ChIP cardiomyocyte_1 GSE85628.TBX5.cardiomyocyte_1 316 bp overlap
TCF12 1 dataset
ChIP A-549 ENCSR000BQQ.TCF12.A-549 221 bp overlap
TEAD3 1 dataset
ChIP HepG2 ENCFF054UUL 214 bp overlap
TFAP2C 2 datasets
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 499 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 254 bp overlap
TSC22D1 1 dataset
ChIP MCF-7 ENCFF552PAC 251 bp overlap
TSC22D2 1 dataset
ChIP MCF-7 ENCFF588BUN 297 bp overlap
ZBTB43 2 datasets
ChIP WTC11 ENCFF058JUB 485 bp overlap
ChIP WTC11 ENCFF058JUB 293 bp overlap
ZC3H4 2 datasets
ChIP K562 ENCFF343JOP 387 bp overlap
ChIP K562 ENCFF343JOP 179 bp overlap
ZFP3 1 dataset
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR581GUY.ZFP3.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 255 bp overlap
ZFX 1 dataset
ChIP MCF-7 GSE102616.ZFX.MCF-7 295 bp overlap
ZKSCAN1 2 datasets
ChIP Hep-G2 ENCSR157CAU.ZKSCAN1.Hep-G2 298 bp overlap
ChIP K-562 ENCSR882ERE.ZKSCAN1.K-562 208 bp overlap
ZNF257 1 dataset
ChIP HEK293T GSE78099.ZNF257.HEK293T 208 bp overlap
ZNF302 1 dataset
ChIP A549 ENCFF333REI 271 bp overlap
ZNF362 2 datasets
ChIP HEK293 ENCSR715QNO.ZNF362.HEK293 129 bp overlap
ChIP HepG2 ENCFF256AZN 149 bp overlap
ZNF776 1 dataset
ChIP HepG2 ENCFF009LSZ 282 bp overlap